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New species of arrowworms (Chaetognatha) from Chazhma Bay (Peter the Great Bay, Sea of Japan).
A. P. Kasatkina, A. F. Sergeev (Pacific Oceanological Institute, Far East Branch, Russian Academy of Sciences, Vladivostok 690041)
Eighteen species of Chaetognatha were discovered in Chazhma Bay (Peter the Great Bay, Sea of Japan). Of these,
4 new species (Leptosagitta ocis sp. n., L. exigua sp. n., L. icis sp. n., and Oligoradiata entis sp. n.) are described as new
for science. Some of specimens had morphological abnormalities, which were not observed in chaetognaths from volcanic bays and active geophysical regions. The ecological situation in Chazhma Bay is analyzed. A list of chaetognaths
along with descriptions and figures of new species are provided. Morphological abnormalities found in all species of
chaetognaths are outlined. (Biologiya Morya, Vladivostok, 2004, vol. 30, no. 6, pp. 440–446).
Key words: arrowworms, Chaetognatha, plankton, morphological abnormalities, radioactive contamination.
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Wlbfheh]by. "Exigua"±f_edZyeZl
;BHEH=BY FHJY lhf 30 ‹ 6 2004
GH<U?<B>U
443
Leptosagitta ocis sp. n.
Jbk:=
>bZ]ghaI_j_^gbcdhg_pieZ\gbdh\ciZjugZqbgZ_lkyihaZ^bmjh\gyaZ^g_]hdhgpZ[jxrgh]h]Zg]eby ;hdh\u_ ieZ\gbdb \ bo i_j_^g_c qZklb ebr_gu
emq_c[_aemq_\ZyahgZbf__lkylhevdh\[hdh\uoieZ\gbdZo Ijhf_`mlhd f_`^m ieZ\gbdZfb c b c iZju
jZ\_g ijhf_`mldm f_`^m [jxrguf ]Zg]eb_f b ieZ\gbdZfbciZju;jxrghc]Zg]ebc^ebgg__lmeh\bsghcqZklbieZ\gbdZciZju
FZl_jbZe =hehlbi Bg\_glZjguc ghf_j /3
H[gZjm`_g\[QZ`fZaZeI_ljZ<_ebdh]hYihgkdh]h
fhjyn_\jZey]\ih\_joghklghfkeh_\h^uijb
hljbpZl_evghcl_fi_jZlmj_
IZjZlbiu Ljb wda_fieyjZ \u^_e_gu ba lhc `_
ijh[u qlh b ]hehlbi Bg\_glZjgu_ ghf_jZ /3
LP-8/3, LP-8/4.
HibkZgb_ ]hehlbiZ L_eh jb]b^gh_ fmkdmebklh_
=heh\Z rbj_ lmeh\bsZ gZ lmeh\bsgho\hklh\hc i_j_]hjh^d_ km`_gby g_l Wda_fieyj gZ c klZ^bb aj_ehklb >ebgZ l_eZ ff >ebgZ o\hklh\h]h hl^_eZ khklZ\ey_lZ[jxrgh]h]Zg]ebyhdhehhl^ebgul_eZ<ujhklulmeh\bsghcdbrdbhlkmlkl\mxlLmeh\bsgZy dbrdZ g_ \Zdmhebabjh\ZgZ g_ kiexs_gZ
\^hjkh\_gljZevghf gZijZ\e_gbb gh g_ ijyfZy dZd m
\k_ohklZevguo\b^h\jh^ZZba\bebklZyF_jpZl_evgZy
i_ley dhjhldZy gZqbgZ_lky kjZam ihaZ^b mjh\gy ]eZa
qmlv \ur_ r_b b jZkiheZ]Z_lky ihjh\gm gZ ]heh\_ b
lmeh\bs_ I_ley bf__l h^gm iZjm keZ[h \ujZ`_gguo
f_^bZevguo \uimdehkl_c I_j_^gbc dhg_p ieZ\gbdh\
c iZju gZqbgZ_lky ihaZ^b mjh\gy aZ^g_]h dhgpZ
[jxrgh]h ]Zg]eby gZ jZkklhygbb \ jZaZ dhjhq_
[jxrgh]h]Zg]ebyIeZ\gbdbciZjudhjhq_ieZ\gbdh\ c iZju ijb[ebabl_evgh \ jZaZ b khklZ\eyxl
ijbf_jghhl^ebgul_eZI_j_^gbcdhg_pieZ\gbdh\ c iZju hlklhbl hl aZ^g_]h dhgpZ ieZ\gbdh\ c
iZjugZjZkklhygb_khklZ\eyxs__hdhehhl^ebgu
l_eZhghjZ\ghjZkklhygbxf_`^m[jxrguf]Zg]eb_f
bieZ\gbdZfbciZju>ebgZieZ\gbdh\ciZjukhklZ\ey_l [he__ hl ^ebgu l_eZ bo lmeh\bsgZy
qZklv dhjhq_ o\hklh\hc \ jZaZ O\hklh\hc ieZ\gbd
aZdjm]e_gguc _]h ^ebgZ gZ l_e_ khklZ\ey_l hl h[s_c^ebgubijb[ebabl_evghhl^ebguo\hklh\h]h
hl^_eZ Emqb g_ihegu_ lhevdh \ [hdh\uo ieZ\gbdZo
i_j_^gyyqZklvdhlhjuoiheghklvxebr_gZemq_c:ev\_heyjgZyldZgvihdju\Z_l\k_l_ehhlr_b^hdZm^Zevgh]hdhgpZmlhesZykvk^hjkZevghcklhjhgugZijhlb\
[jxrgh]h ]Zg]eby b gZ lmeh\bs_ f_`^m ieZ\gbdZfb
1cbciZjuGZ]heh\_h^gZiZjZjy^h\s_lbghd^\_
iZju jy^h\ am[qbdh\ S_lbghd ih i_j_^gbo am[qbdh\ihaZ^gboam[qbdh\ih\dZ`^hfjy^m=eZaZk
l_fgufijyfhm]hevgufib]f_glhf\p_gljZevghcqZklb K_f_ggu_ f_rhqdb m^ebg_ggu_ khklZ\eyxl hdheh
hl ^ebgu l_eZ b hl ^ebgu o\hklh\h]h hl^_eZ
hgbkhijbdZkZxlkydZdk[hdh\ufblZdbko\hklh\uf
ieZ\gbdhf Ybqgbdb ^hklb]Zxl mjh\gy i_j_^g_]h dhgpZieZ\gbdh\ciZju
;BHEH=BY FHJY lhf 30 ‹ 6 2004
Jbk =hehlbiu : = ± Leptosagitta ocis sp. n : ± h[sbc
\b^=±]eZa; < – Oligoradiata entis sp. n;±h[sbc\b^
<±]eZa
D:K:LDBG:K?J=??<
444
Baf_gqb\hklv >ebgZ iheh\haj_euo hkh[_c dhe_[e_lky hl ^h ff Z hlghkbl_evgZy ^ebgZ o\hklh\h]h hl^_eZ fh`_l khklZ\eylv ± hl ^ebgu
l_eZ[jxrgh]h]Zg]eby±±hl^ebgul_eZ
>bnn_j_gpbZevguc ^bZ]gha <b^ [ebahd d L. exigua VSQghhlebqZ_lkyhlg_]hjy^hfijbagZdh\kj_^bgbogZb[he__aZf_lgu_gZebqb_[_aemq_\uoahgm
L. ocis VSQjZkiheh`_gb_ieZ\gbdh\m L. ocis sp. n.
ijhf_`mlhd f_`^m ieZ\gbdZfb c b c iZju jZ\_g
jZkklhygbx f_`^m [jxrguf ]Zg]eb_f b ieZ\gbdZfb
1ciZjuZmL. exigua VSQhgh^ebgg__[jxrghc
]Zg]ebc m L. exigua VS Q dhjhq_ lmeh\bsghc qZklb
ieZ\gbdh\ciZjumL. ocis VSQ±^ebgg__
K\_^_gbyih[bheh]bbIhkhklhygbx]hgZ^fh`gh dhk\_ggh km^blv hjZafgh`_gbb hgh ijhbkoh^bl
hq_\b^gh ijb hljbpZl_evghc l_fi_jZlmj_ \^_dZ[j_
Iheh\haj_euowda_fieyjh\\_kghcg_hlf_q_gh
Wlbfheh]by. "Ocis"±[ukljuceZl
Oligoradiata entis sp. n.
Jbk;<
>bZ]gha K_f_ggu_ f_rhqdb m^Ze_gu dZd hl [hdh\uolZdbhlo\hklh\h]hieZ\gbdZgZjZ\gh_jZkklhygb_ Lmeh\bsgZy qZklv ieZ\gbdh\ c iZju agZqbl_evghdhjhq_boo\hklh\hcqZklbijbf_jgh\jZaZ
FZl_jbZe=hehlbiBg\_glZjgucghf_j2/5
H[gZjm`_g \ [QZ`fZ aZe I_ljZ <_ebdh]h Yihgkdh]h
fhjyn_\jZey]
IZjZlbiu >\Z wda_fieyjZ \u^_e_gu ba lhc `_
ijh[u qlh b ]hehlbi Bg\_glZjgu_ ghf_jZ 2/5
OLR-1/14.
HibkZgb_]hehlbiZL_eh^h\hevgh\yeh_ijh^hev
gu_ fmkdmevgu_ ly`b rbjhdb_ ohjhrh ijhk\_qb\Zxl
=heh\Z rbj_ lmeh\bsZ gZ lmeh\bsgho\hklh\hc i_j_]hjh^d_km`_gbyg_lWda_fieyjgZcklZ^bbaj_ehklb
>ebgZl_eZff>ebgZo\hklh\h]hhl^_eZkhklZ\ey_l Z[jxrgh]h ]Zg]eby hdheh hl ^ebgu
l_eZDbr_qgu_\ujhklugZkj_^g_cdbrd_g_[hevrb_
iehlghijb`Zluddbrd_Lmeh\bsgZydbrdZg_\Zdmhebabjh\ZgZ F_jpZl_evgZy i_ley dhjhldZy gZqbgZ_lky
ihaZ^bmjh\gy]eZaaZdjm]ey_lkygZ]heh\_I_leybf__l lhevdh h^gm iZjm keZ[h \ujZ`_gguo f_^bZevguo
\uimdehkl_c I_j_^gbc dhg_p ieZ\gbdh\ c iZju gZqbgZ_lkyihaZ^bmjh\gyaZ^g_]hdhgpZ[jxrgh]h]Zg]eby gZ jZkklhygbb dhlhjh_ \ jZaZ dhjhq_ ^ebgu
[jxrgh]h]Zg]ebyqlhkhklZ\ey_lhl^ebgul_eZ
IeZ\gbdbciZjudhjhq_ieZ\gbdh\ciZjuijb[ebabl_evgh \ jZaZ bo ^ebgZ ^hklb]Z_l ijbf_jgh hl^ebgul_eZI_j_^gbcdhg_pieZ\gbdh\ciZjuhlklhblhlaZ^g_]hdhgpZieZ\gbdh\ciZjugZjZkklhygb_ khklZ\eyxs__ hl ^ebgu l_eZ hgh [hevr_
jZkklhygby f_`^m [jxrguf ]Zg]eb_f b ieZ\gbdZfb
1c iZju \ jZaZ >ebgZ ieZ\gbdh\ c iZju [he__
hl^ebgul_eZbolmeh\bsgZyqZklvdhjhq_o\hklh\hc ijb[ebabl_evgh \ jZaZ O\hklh\hc ieZ\gbd
aZdjm]e_gguc _]h ^ebgZ gZ l_e_ khklZ\ey_l hl
^ebgu l_eZ b hdheh hl ^ebgu o\hklh\h]h hl^_eZ
Emq_cg_l g_ lhevdh \ [hdh\uo ieZ\gbdZo gh b \o\h-
klh\hf hl^_evgu_ j_kgbpu ihy\eyxlky ebrv m iheh\haj_euo `b\hlguo :ev\_heyjgZy ldZgv hlkmlkl\m_l
GZ ]heh\_ h^gZ iZjZ jy^h\ s_lbghd b ^\_ iZju jy^h\
am[qbdh\ S_lbghd ih i_j_^gbo am[qbdh\ ih ±
aZ^gboam[qbdh\ih±\dZ`^hfjy^m =eZaZ k l_fguf ijyfhm]hevguf ib]f_glguf iylghf \ p_gljZevghcqZklbK_f_ggu_f_rhqdbm^ebg_ggu_khklZ\eyxl
hl^ebgul_eZbhdhehhl^ebguo\hklh\h]h
hl^_eZ F_rhqdb m^Ze_gu dZd hl [hdh\uo lZd b hl
o\hklh\h]h ieZ\gbdZ gZ jZ\gh_ jZkklhygb_ khklZ\eyxs__ hl ^ebgu l_eZ b hdheh hl ^ebgu
o\hklh\h]h hl^_eZ Ybqgbdb khklZ\eyxl hl ^ebgu l_eZ hgb g_ ^hklb]Zxl mjh\gy i_j_^g_]h dhgpZ
ieZ\gbdh\ciZju
Bg^b\b^mZevgZybaf_gqb\hklv Dhebq_kl\h emq_c
\ ieZ\gbdZo c iZju b o\hklh\hf ieZ\gbd_ fh`_l
\Zjvbjh\Zlvhl^h
>bnn_j_gpbZevguc^bZ]ghaHlebqZ_lkyhlO. mitis .DVVDWNLQD jy^hf ijbagZdh\ gZb[he__ aZf_lgu_mO. mitislmeh\bsgZyqZklvieZ\gbdh\ciZju
ijb[ebabl_evghjZ\gZbebg_fgh]hdhjhq_boo\hklh\hc qZklb Z m O. entis VS Q ± agZqbl_evgh dhjhq_ bo
o\hklh\hc qZklb ijbf_jgh \ jZaZ m O. mitis
bf__lkyZev\_heyjgZyldZgvZk_f_ggu_f_rhqdbijb[eb`_gu d o\hklh\hfm ieZ\gbdm O. entis VS Q bf__l
koh^kl\hk O. pellucida.DVVDWNLQDHW6HOLYDQRYDgh
hlebqZ_lky hl wlh]h \b^Z jy^hf ijbagZdh\ gZb[he__
aZf_lgu_badhlhjuok_f_ggu_f_rhqdbmO. pellucidakhijbdZkZxlkyk[hdh\ufbghm^Ze_guhlo\hklh\h]hieZ\gbdZi_j_^gbcdhg_pieZ\gbdh\ciZjum
O. pellucida gZqbgZ_lky ihaZ^b mjh\gy aZ^g_]h dhgpZ
[jxrgh]h ]Zg]eby gZ jZkklhygbb \ jZaZ dhjhq_
[jxrgh]h]Zg]eby
K\_^_gbyih[bheh]bbIhkhklhygbx]hgZ^fh`ghdhk\_gghkm^blvhjZafgh`_gbbhq_\b^ghhghijhbkoh^blijbhljbpZl_evghcl_fi_jZlmj_\^_dZ[j_Iheh\haj_euowda_fieyjh\\_kghcg_hlf_q_gh
Wlbfheh]by. "Entis"±ijhk\_qb\ZxsZyeZl
HibkZgb_fhjnheh]bq_kdbohldehg_gbc
mfhjkdboklj_ehd\[QZ`fZ
G_h[oh^bfh hlf_lblv qlh i_j_qbke_ggu_ gb`_
fhjnheh]bq_kdb_ hldehg_gby g_ y\eyxlky f_oZgbq_kdbfb ih\j_`^_gbyfb Fgh]he_lgbc hiul jZ[hlu Z\lhjh\ klZlvb m[_^bl_evgh ihdZaZe qlh f_oZgbq_kdb_
ih\j_`^_gby m fhjkdbo klj_ehd fh]ml ihy\blvky \ j_amevlZl_ g_ hl[hjZ Z jZa[hjZ ijh[ g_hiulgufb ex^vfb dhlhju_ ihevamykv ibgp_lhf k ijyfufb dhgpZfb
kbevghk`bfZxl`b\hlgh_ihi_j_dlmeh\bsZFujZa[bjZeb ijh[u ibgp_lhf k aZ]gmlufb dhgpZfb beb k
ihfhsvxibi_ldbIjbf_qZl_evghqlhbmZghfZevguo
hkh[_c hlh[jZgguo ba [ QZ`fZ b m hkh[_c ba ]_hnbabq_kdb Zdlb\guo j_]bhgh\ hklZ_lky g_ih\j_`^_gguf
gZb[he__g_`gucbmya\bfuchj]Zgfhjkdboklj_ehd±
f_jpZl_evgZy i_ley dhlhjZy mljZqb\Z_lky ijb g_ZddmjZlghff_oZgbq_kdhfi_j_f_s_gbbwda_fieyjh\
Gb \ h^ghf ba j_]bhgh\ h[jZ[hlZgu lukyqb
ijh[±hl:leZglbdb^h<hklhqghDblZckdh]hfhjylZ;BHEH=BY FHJY lhf 30 ‹ 6 2004
GH<U?<B>U
dbo fhjnheh]bq_kdbo hldehg_gbc m fhjkdbo klj_ehd
dZd\[QZ`fZg_\klj_qZehkvH^gZdhmfhjkdboklj_ehd ba [QZ`fZ g_h[gZjm`_gufhjnheh]bq_kdb_ ZghfZebboZjZdl_jgu_^ey`b\hlguoh[blZxsbo\\h^Zo
\medZgbq_kdbo[molbk_ckfbq_kdbZdlb\guoj_]bhgh\
baf_g_gby \ ib]f_glghf gmde_Zjghf y^j_ Z bgh]^Z b
ihegh_ bkq_agh\_gb_ ]eZaZ Z lZd`_ ebabk furp DZkZldbgZ.DVVDWNLQDHWDO
GZb[he__ qZklh \klj_qZxsb_ky ZghfZevgu_ hldehg_gbymfhjkdboklj_ehd
JZajmr_gb_ ieZ\gbdh\hc ieZklbgdb ih djZyf
f_`^memq_cIjbq_femqbhklZxlkyg_ih\j_`^_ggufb
bqZs_\k_]hbodjZylhjqZlaZij_^_eZfbieZ\gbdh\uo
ieZklbghd jbk : Bgh]^Z \ j_amevlZl_ jZajmr_gby
ieZklbgdbdjZyieZ\gbdh\g_bf_xl[ZojhfqZlhcdZ_fdbbak\h[h^guoemq_cdZdgZjbk:Z\u]ey^yldZd
i_lmrbgu_]j_[gbjbk;G_h[oh^bfhhlf_lblvqlh
ijbf_oZgbq_kdhfih\j_`^_gbbjZajmrZ_lkyg_lhevdh
djZ_\Zyf_f[jZgZZbgh]^Zb\kyieZ\gbdh\ZyieZklbgdZghih\j_`^ZxlkybieZ\gbdh\u_emqbihwlhfmih^h[ghc dZjlbgu mf_oZgbq_kdb ih\j_`^_gguo hkh[_c
g_ gZ[ex^Z_lky >ey ZghfZevguo wda_fieyjh\ ba [
QZ`fZi_lmrbguc]j_[_gv±wlhbooZjZdl_jgucijbagZd Ijbf_qZl_evgh qlh m ihjZ`_gguo wda_fieyjh\
445
\k_o\b^h\bf_xsboijZ\bevgu_]mklu_emqbhgbkhojZgyxlky \h \k_o ieZ\gbdZo ihwlhfm fu kqblZ_f
ieZ\gbdh\u_ emqb lZdkhghfbq_kdbf ijbagZdhf b bo
hlkmlkl\b_g_fh`_lkem`blvijbagZdhfZghfZevghklb
Hq_\b^ghwlhk\yaZghkl_fqlhemqbkhklhylba[he__
ijhqgh]h obfbq_kdh]h \_s_kl\Z q_f ieZklbgdZ ieZ\gbdZ'XYHU6DODWLZdb_hldehg_gby\klj_qZxlky m\b^h\ h[eZ^Zxsbo ]mklufb fgh]hqbke_ggufb
emqZfb\ieZ\gbdZoLeptosagitta ocis sp. n.; L. exigua sp.
n.; L. icis sp. n.; L. nudata Kassatkina, 1973; L. collariata
Kassatkina, 1973; L. uschakovi Kassatkina, 1973; L. eris
Kassatkina et Selivanova, 2003; Aidanosagitta scarlatoi
Kassatkina, 1971; A. macilenta Kassatkina, 1971; A. acus
Kassatkina et Selivanova, 2003; A. pilum Kassatkina et
Selivanova, 2003; A. murex Kassatkina et Selivanova, 2003;
A. bella Kassatkina et Selivanova, 2003; A. venusta Kassatkina et Selivanova, 2003; Parasagitta septicoela Kassatkina,
JZajmr_gb_ djZ_\hc f_f[jZgu m \b^h\ g_ bf_xsbo ]mkluo emq_c \ ieZ\gbdZo ijb\h^bl d lhfm qlh
rbjhdbc ieZ\gbd \u]ey^bl dZd madZy ihehkdZ beb kh\k_fbkq_aZ_lWlh\klj_qZ_lkym\b^h\bajh^Z2OigoradiaWDZlZd`_mL. alba Kassatkina et Selivanova, 2003.
JZajmr_gb_ bklhgq_gb_ [ZaZevghc f_f[jZgu
<wlhfkemqZ_s_lbgdbbam[qbdbhl^_eyxlkyhl]heh-
JbkFhjnheh]bq_kdb_ZghfZebbmkZ]bll:±o\hklh\hcieZ\gbd(L. exigua sp. n.);±qZklv[hdh\h]hieZ\gbdZkih\j_`^_gghc[ZaZevghcieZklbgdhc(L. icis sp. n.)<±]heh\ZbqZklvlmeh\bsZgZ]heh\_iheghklvxhlkmlkl\m_lam[gh_\hhjm`_gb_ZmqZklhddbrdb
bf__lih\j_`^_gby(P. septicoela)=±]heh\ZbqZklvlmeh\bsZgZ]heh\_iheghklvxkhojZgy_lkyam[gh_\hhjm`_gb_ZmqZklhddbrdb
bf__lih\j_`^_gby(L. exigua sp. n.)> ±qZklvlmeh\bsgh]hhl^_eZkZghfZevgufmqZkldhfZev\_heyjghcldZgb(L. icis sp. n.).
;BHEH=BY FHJY lhf 30 ‹ 6 2004
D:K:LDBG:K?J=??<
446
\uBgh]^Zhgbkh\k_fhllhj]Zxlky\lZdhfkemqZ_gZ[ex^Z_lky kljZggZy ^ey ij_^klZ\bl_e_c lbiZ &KDHWR
gnaWKDdZjlbgZeukZy]heh\Z[_aam[gh]h\hhjm`_gby
jbk<Ijbf_qZl_evghqlhfurpu]heh\uijbwlhf
khojZgyxlky \ hlebqb_ gZijbf_j hl wda_fieyjh\ ba
]_hnbabq_kdb Zdlb\gh]h jZchgZ [ DjZl_jghc LZdb_
ZghfZevgu_ wda_fieyju h[gZjm`_gu kj_^b jh^h\ Leptosagitta (L. exigua sp. n., L. icis sp. n.) b Parasagitta
(P. septicoela).
JZajmr_gb_ dbr_qgh]h wibl_eby jbk < =
Ba\_klgh qlh mfhjkdbo klj_ehd \ jZchgZo k \ukhdhc
]_hnbabq_kdhc Zdlb\ghklvx iheghklvx khojZgyxlky
ldZgb dbr_qgh]h b ihdjh\gh]h wibl_eby DZkZldbgZ
JZajmr_gb_ dbr_qgh]h wibl_eby m fhjkdbo
klj_ehdg_hlf_q_ghgb\h^ghfbawdheh]bq_kdbg_[eZ]hihemqguo j_]bhgh\ F_oZgbq_kdb_ ih\j_`^_gby
dbrdb ijb jZa[hj_ ijh[u \hafh`gu ebrv \ lhf kemqZ__kebkbevghk`Zlvibgp_lhfl_eh`b\hlgh]hh^gZdh ijb wlhf ih\j_`^Zxlky b gZjm`gu_ ldZgb <
[QZ`fZh[gZjm`_guwda_fieyjumdhlhjuoijbiheghc khojZgghklb gZjm`guo ldZg_c [ue ih\j_`^_g dbr_qgbd(P. septicoela, L. exigua sp. n.).
JZajmr_gb_ Zev\_heyjghc ldZgb qZklv de_lhd
dhlhjhc hl^_ey_lky ijb wlhf djZy Zev\_heyjghc h[hehqdb klZgh\ylky g_ jh\gufb Z dZd [u bat_^_ggufb
jbk>LZdb_hldehg_gby\klj_qZxlkym\b^h\h[eZ^ZxsboohjhrhjZa\blhcZev\_heyjghcldZgvxLeptosagitta ocis sp. n.; L. exigua sp. n.; L. icis sp. n.; L. nudata
Kassatkina, 1973; L. collariata Kassatkina, 1973;
L. uschakovi Kassatkina, 1973; L. eris Kassatkina et
Selivanova, 2003; Aidanosagitta scarlatoi Kassatkina,
1971; A. macilenta Kassatkina, 1971; A. acus Kassatkina
et Selivanova, 2003; A. pilum Kassatkina et Selivanova,
2003; A. murex Kassatkina et Selivanova, 2003; A. bella
Kassatkina et Selivanova, 2003; A. venusta Kassatkina et
Selivanova, 2003.
:\lhju [eZ]h^Zjyl <: =hjyq_\Z <D KZ\q_gdh
bH<IhghfZj_\ZaZihfhsvijbhl[hj_ijh[
KIBKHDEBL?J:LMJU
:klZoh\ :K Eblheh]by hkZ^dh\ fZl_jbdh\hc hdjZbgu \hklhdZ:abb<eZ^b\hklhd>ZevgZmdZk
;_eZg L: Wdheh]bq_kdb_ bkke_^h\Zgby \ aZeb\_ Klj_ehd b
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Kassatkina A.P., Pantelyev E.V., Yurasov G.I. et al. MorphologiFDODQRPDOLHVRIKKDHWRJQDWKDLQ3HWHUWKH*UHDW%D\RIWKH
Sea of Japan // Oceanography of the Japan Sea. Vladivostok: Dalnauka. 2001. P. 255–259.
Sergeev A.F., Goryachev V.A., Soyfer V.N. et al. The oceanography aspect of radiation situation in Chazhma Bay of Japan
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;BHEH=BY FHJY lhf 30 ‹ 6 2004
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