ɘɥɩɦɩɞɣɺ ɫɛɬɭɠɨɣɤ Ecology of plants əɞ ɋɩɬɬɣɣ: ɸɥɩɦɩɞɣɺ, ɫɛɢɝɣɭɣɠʌ 4, 2011 The South of Russia: ecology, development. ʋ4, 2011 ɘɅɉɆɉȾɃɚ ɋȻɌɍɀɈɃɄ ɍȾɄ: 582.734.3.02 ȽɆɃɚɈɃɀ ɘɅɉɆɉȾɃɒɀɌɅɃɐ ɏȻɅɍɉɋɉȽ ɈȻ ɈȻɅɉɊɆɀɈɃɀ ȼɃɉɆɉȾɃɒɀɌɅɃ ȻɅɍɃȽɈɖɐ ȽɀɔɀɌɍȽȽɊɆɉȿȻɐ CRATAEGUS PENTAGYNA WALDST ET KIT Ƚ ɎɌɆɉȽɃɚɐ ȿȻȾɀɌɍȻɈȻ 2011 ɈɦɚɪɢɟɜɚɅȼ Ⱦɚɝɟɫɬɚɧɫɤɢɣɝɨɫɭɞɚɪɫɬɜɟɧɧɵɣɭɧɢɜɟɪɫɢɬɟɬ ɂɡɭɱɟɧɯɢɦɢɱɟɫɤɢɣɫɨɫɬɚɜɩɥɨɞɨɜ&UDWDHJXVSHQWDJ\QD:DOGVWHW.LWɜɡɚɜɢɫɢɦɨɫɬɢɨɬɦɟɫɬɨɩɪɨɢɡɪɚɫɬɚɧɢɹȼɵɹɜɥɟɧɚ ɢɡɦɟɧɱɢɜɨɫɬɶ ɫɨɫɬɚɜɚ ɩɥɨɞɨɜ ɨɬ ɞɟɣɫɬɜɢɹ ɮɚɤɬɨɪɨɜ ɨɤɪɭɠɚɸɳɟɣ ɫɪɟɞɵ. The chemical composition of the fruits of Crataegus pentagyna Waldst et Kit against the habitat is studied. The variability of the fruits of the influence of environmental factors. Ʉɥɸɱɟɜɵɟɫɥɨɜɚ ɛɨɹɪɵɲɧɢɤɛɢɨɥɨɝɢɱɟɫɤɢɚɤɬɢɜɧɵɟɜɟɳɟɫɬɜɚɷɤɨɥɨɝɢɱɟɫɤɢɟɮɚɤɬɨɪɵ. Key words: Hawthorne, biologically active substances, environmental factors. Ȼɨɹɪɵɲɧɢɤ ± ɨɞɢɧ ɢɡ ɞɪɟɜɧɟɣɲɢɯ ɥɟɤɚɪɫɬɜɟɧɧɵɯ ɪɚɫɬɟɧɢɣ ɢɫɩɨɥɶɡɭɟɦɵɯ ɜ ɦɟɞɢɰɢɧɟ ɫɨ ɜɪɟɦɟɧȾɢɨɫɤɨɪɢɞɚ,ɜɧɷ. ɂɫɩɨɥɶɡɨɜɚɧɢɟ ɛɨɹɪɵɲɧɢɤɚ ɜɦɟɞɢɰɢɧɟ ɫɜɹɡɚɧɨ ɫ ɲɢɪɨɤɢɦ ɫɩɟɤɬɪɨɦ ɟɝɨ ɞɟɣɫɬɜɢɹ ɧɚ ɨɪɝɚ ɧɢɡɦ Ɉɧ ɫɧɢɠɚɟɬ ɜɨɡɛɭɞɢɦɨɫɬɶ ɰɟɧɬɪɚɥɶɧɨɣ ɧɟɪɜɧɨɣ ɫɢɫɬɟɦɵ ɬɨɧɢɡɢɪɭɟɬ ɫɟɪɞɟɱɧɨ-ɫɨɫɭɞɢɫɬɭɸ ɫɢɫɬɟɦɭ ɭɫɢɥɢɜɚɟɬ ɦɨɡɝɨɜɨɟ ɤɪɨɜɨɨɛɪɚɳɟɧɢɟ ɫɧɢɠɚɟɬ ɤɪɨɜɹɧɨɟ ɞɚɜɥɟɧɢɟ ɨɛɥɚɞɚɟɬ ɛɚɤɬɟɪɢɰɢɞ ɧɵɦɢɫɜɨɣɫɬɜɚɦɢɢɩɪɗɬɨɛɟɡɭɫɥɨɜɧɨɫɜɹɡɚɧɨɫɨɫɨɛɟɧɧɨɫɬɹɦɢɯɢɦɢɱɟɫɤɨɝɨɫɨɫɬɚɜɚɩɥɨɞɨɜɛɨɹ ɪɵɲɧɢɤɚɹɜɥɹɸɳɢɯɫɹɛɨɝɚɬɵɦɢɫɬɨɱɧɢɤɨɦɛɢɨɥɨɝɢɱɟɫɤɢɚɤɬɢɜɧɵɯɜɟɳɟɫɬɜȻȺȼɉɨɷɬɨɦɭɩɥɨ ɞɵɛɨɹɪɵɲɧɢɤɚɦɨɠɧɨɪɚɫɫɦɚɬɪɢɜɚɬɶɤɚɤɜɟɫɶɦɚɩɟɪɫɩɟɤɬɢɜɧɨɟɫɵɪɶɟɞɥɹɢɫɩɨɥɶɡɨɜɚɧɢɹɜɮɚɪɦɚ ɰɢɢɢɩɢɳɟɜɨɣɩɪɨɦɵɲɥɟɧɧɨɫɬɢ>]. ɂɡɭɱɟɧɢɟɜɥɢɹɧɢɹɷɤɨɥɨɝɢɱɟɫɤɢɯɮɚɤɬɨɪɨɜɢɭɫɥɨɜɢɣɦɟɫɬɨɨɛɢɬɚɧɢɹɧɚɫɨɞɟɪɠɚɧɢɟɜɪɚɫɬɟ ɧɢɹɯɛɢɨɥɨɝɢɱɟɫɤɢɚɤɬɢɜɧɵɯɜɟɳɟɫɬɜɱɪɟɡɜɵɱɚɣɧɨɜɚɠɧɨɤɚɤɞɥɹɪɚɡɪɚɛɨɬɤɢɬɟɨɪɟɬɢɱɟɫɤɢɯɜɨɩɪɨ ɫɨɜ ɪɟɫɭɪɫɨɜɟɞɟɧɢɹ ɬɚɤ ɢ ɞɥɹ ɨɛɟɫɩɟɱɟɧɢɹ ɦɟɞɢɰɢɧɫɤɨɣ ɩɪɨɦɵɲɥɟɧɧɨɫɬɢ ɜɵɫɨɤɨɤɚɱɟɫɬɜɟɧɧɵɦ ɫɵɪɶɟɦ ɗɬɨ ɩɨɡɜɨɥɹɟɬ ɜɵɹɜɢɬɶ ɨɩɬɢɦɚɥɶɧɵɟ ɪɚɣɨɧɵ ɡɚɝɨɬɨɜɤɢ ɞɥɹ ɤɚɠɞɨɝɨɜɢɞɚ ɢ ɡɚɝɨɬɚɜɥɢɜɚɬɶ ɫɵɪɶɟɜɵɫɨɤɨɝɨɤɚɱɟɫɬɜɚ Ɉɬɞɟɥɶɧɵɟɧɚɛɥɸɞɟɧɢɹɧɚɞɢɡɦɟɧɱɢɜɨɫɬɶɸɯɢɦɢɱɟɫɤɨɝɨɫɨɫɬɚɜɚɪɚɫɬɟɧɢɣɢɟɟɡɚɜɢɫɢɦɨɫɬɶɸ ɨɬɤɥɢɦɚɬɚɩɪɨɢɡɜɨɞɢɥɢɫɶɧɟɨɞɧɨɤɪɚɬɧɨɨɞɧɚɤɨɜɵɜɨɞɵɢɡɷɬɢɯɧɚɛɥɸɞɟɧɢɣɞɟɥɚɥɢɫɶɥɢɲɶɜɫɚ ɦɨɣɨɛɳɟɣɮɨɪɦɟ>]. ɇɚɯɢɦɢɱɟɫɤɢɣɫɨɫɬɚɜɥɟɤɚɪɫɬɜɟɧɧɵɯɪɚɫɬɟɧɢɣɛɨɥɶɲɨɟɜɥɢɹɧɢɟɨɤɚɡɵɜɚɸɬɪɚɡɥɢɱɧɵɟɮɚɤɬɨɪɵ ɨɤɪɭɠɚɸɳɟɣ ɫɪɟɞɵ ɝɟɨɝɪɚɮɢɱɟɫɤɢɣ ɤɥɢɦɚɬɢɱɟɫɤɢɣ ɷɞɚɮɢɱɟɫɤɢɣ ɨɪɨɝɪɚɮɢɱɟɫɤɢɣ ɛɢɨɬɢɱɟɫɤɢɣ ɂɦɟɧɧɨ ɫɨɜɨɤɭɩɧɨɫɬɶ ɜɥɢɹɧɢɹ ɤɚɤ ɨɧɬɨɝɟɧɟɬɢɱɟɫɤɢɯ ɮɚɤɬɨɪɨɜ, ɬɚɤ ɢ ɮɚɤɬɨɪɨɜ ɨɤɪɭɠɚɸɳɟɣ ɫɪɟɞɵ, ɢ ɨɩɪɟɞɟɥɹɟɬɜɤɨɧɟɱɧɨɦɫɱɟɬɟɯɢɦɢɱɟɫɤɢɣɫɨɫɬɚɜɥɟɤɚɪɫɬɜɟɧɧɵɯɪɚɫɬɟɧɢɣɢɟɝɨɢɡɦɟɧɱɢɜɨɫɬɶ>]. ɇɚɦɢɛɵɥɚ ɪɚɫɫɦɨɬɪɟɧɚɡɚɜɢɫɢɦɨɫɬɶɫɨɞɟɪɠɚɧɢɹ ɛɢɨɥɨɝɢɱɟɫɤɢɚɤɬɢɜɧɵɯ ɜɟɳɟɫɬɜ ɨɬɷɤɨɥɨ ɝɨ-ɝɟɨɝɪɚɮɢɱɟɫɤɢɯɮɚɤɬɨɪɨɜɬɚɤɢɯɤɚɤɫɪɟɞɧɹɹɬɟɦɩɟɪɚɬɭɪɚɢɫɭɦɦɚɨɫɚɞɤɨɜɡɚɜɟɝɟɬɚɰɢɨɧɧɵɣɩɟ ɪɢɨɞɛɨɹɪɵɲɧɢɤɚɜɵɫɨɬɚɧɚɞɭɪɨɜɧɟɦɦɨɪɹ Ⱦɥɹɜɵɹɜɥɟɧɢɹɜɥɢɹɧɢɹɬɚɤɢɯɮɚɤɬɨɪɨɜɤɚɤɬɟɦɩɟɪɚɬɭɪɚɢɜɥɚɠɧɨɫɬɶɜɨɡɞɭɯɚɦɵ ɫɨɛɢɪɚɥɢ ɨɛɪɚɡɰɵɩɥɨɞɨɜɜɨɞɧɨɦɢɬɨɦɠɟɦɟɫɬɨɨɛɢɬɚɧɢɢɢɜɨɞɧɭɢɬɭɠɟɮɚɡɭɩɨɥɧɨɝɨɩɥɨɞɨɧɨɲɟɧɢɹɜ ɬɟɱɟɧɢɟɧɟɫɤɨɥɶɤɢɯɥɟɬɌɟɦɩɟɪɚɬɭɪɧɵɟɞɚɧɧɵɟɚɬɚɤɠɟɞɚɧɧɵɟɨɤɨɥɢɱɟɫɬɜɟɨɫɚɞɤɨɜɛɵɥɢɩɨɥɭ ɱɟɧɵɩɨɥɢɬɟɪɚɬɭɪɧɵɦɢ ɤɚɪɬɨɝɪɚɮɢɱɟɫɤɢɦɞɚɧɧɵɦ>1, 3@ɢɜɊɟɫɩɭɛɥɢɤɚɧɫɤɨɦɝɢɞɪɨɦɟɬɟɨɪɨɥɨɝɢ ɱɟɫɤɨɦɰɟɧɬɪɟɊɟɫɩɭɛɥɢɤɢȾɚɝɟɫɬɚ ɩɨɲɟɫɬɢɫɬɚɧɰɢɹɦ. Ɇɟɬɟɨɪɨɥɨɝɢɱɟɫɤɢɟ ɞɚɧɧɵɟ ɩɨ ɩɭɧɤɬɚɦ ɝɞɟ ɨɬɫɭɬɫɬɜɭɸɬ ɫɬɚɧɰɢɢ ɩɪɢɪɚɜɧɢɜɚɥɢ ɤ ɛɥɢɡɥɟɠɚ ɳɢɦɫɬɚɧɰɢɹɦɫȻɟɥɢɞɠɢɫȼɟɥɢɤɟɧɬɫɄɭɥɟɝɭ ɫɏɭɱɧɢɤɫɬɚɧɰɢɢȾɟɪɛɟɧɬɫɗɪɩɟɥɢ± ɤɫɬɚɧɰɢɢ ȻɭɣɧɚɤɫɤɫɍɱɤɟɧɬɩɄɪɢɜɚɹɛɚɥɤɚ± ɤɫɬɚɧɰɢɢɆɚɯɚɱɤɚɥɚɫȽɚɦɢɹɯ± ɤɫɬɚɧɰɢɢɏɚɫɚɜɸɪɬ ɏɢɦɢɱɟɫɤɢɣ ɫɨɫɬɚɜ ɩɥɨɞɨɜ ɛɨɹɪɵɲɧɢɤɚ ɩɹɬɢɩɟɫɬɢɱɧɨɝɨ ɢɫɫɥɟɞɨɜɚɥɢ ɩɨ ɧɢɠɟ ɩɪɟɞɫɬɚɜɥɟɧ ɧɵɦɦɟɬɨɞɢɤɚɦ Ⱦɥɹ ɤɨɥɢɱɟɫɬɜɟɧɧɨɝɨɨɩɪɟɞɟɥɟɧɢɹ ɫɭɦɦɵ ɮɥɚɜɨɧɨɢɞɨɜɢɫɩɨɥɶɡɨɜɚɥɚɫɶɦɟɬɨɞɢɤɚɞɢɮɮɟɪɟɧ ɰɢɚɥɶɧɨɣ ɫɩɟɤɬɪɨɮɨɬɨɦɟɬɪɢɢ ɨɫɧɨɜɚɧɧɚɹ ɧɚ ɪɟɚɤɰɢɢ ɤɨɦɩɥɟɤɫɨɨɛɪɚɡɨɜɚɧɢɹ ɫ ɪɚɫɬɜɨɪɨɦ ɚɥɸɦɢ 28 ɘɥɩɦɩɞɣɺ ɫɛɬɭɠɨɣɤ Ecology of plants əɞ ɋɩɬɬɣɣ: ɸɥɩɦɩɞɣɺ, ɫɛɢɝɣɭɣɠʌ 4, 2011 The South of Russia: ecology, development. ʋ4, 2011 ɧɢɹɯɥɨɪɢɞɚɜɫɥɚɛɨɤɢɫɥɨɣɫɪɟɞɟ>@ȼɭɫɥɨɜɢɹɯɦɟɬɨɞɢɤɢɪɟɚɤɰɢɹɹɜɥɹɟɬɫɹɫɟɥɟɤɬɢɜɧɨɣɞɥɹɮɥɚ ɜɨɧɨɢɞɨɜɢɩɨɡɜɨɥɹɟɬɨɩɪɟɞɟɥɢɬɶɢɯɫɨɞɟɪɠɚɧɢɟɜɩɟɪɟɫɱɟɬɟɧɚɪɭɬɢɧɢɢɥɢɤɜɟɪɰɟɬɢɧɜɩɪɢɫɭɬ ɫɬɜɢɢɞɪɭɝɢɯɜɟɳɟɫɬɜ Ʉɨɥɢɱɟɫɬɜɟɧɧɨɟɨɩɪɟɞɟɥɟɧɢɟɤɢɫɥɨɬɵɚɫɤɨɪɛɢɧɨɜɨɣɩɪɨɜɨɞɢɥɢɮɨɬɨɤɨɥɨɪɢɦɟɬɪɢɱɟɫɤɢɦɦɟ ɬɨɞɨɦɨɫɧɨɜɚɧɧɨɦɧɚɪɟɚɤɰɢɢɨɤɢɫɥɟɧɢɹɤɢɫɥɨɬɵɚɫɤɨɪɛɢɧɨɜɨɣɠɟɥɬɵɦɮɨɫɮɨɪɧɨ-ɦɨɥɢɛɞɟɧɨɜɵɦ ɝɟɬɟɪɨɰɢɤɥɨɦɫɩɨɫɥɟɞɭɸɳɢɦɨɩɪɟɞɟɥɟɧɢɟɦɩɨɝɥɨɳɟɧɢɹɮɨɫɮɨɪɧɨ-ɦɨɥɢɛɞɟɧɨɜɨɝɨɤɨɦɩɥɟɤɫɚɩɪɢ Ȝ ɧɦ[12]. ɋɨɞɟɪɠɚɧɢɟɚɧɬɨɰɢɚɧɨɜ ɨɩɪɟɞɟɥɹɥɢ ɫɩɟɤɬɪɨɮɨɬɨɦɟɬɪɢɱɟɫɤɢ ɩɨ ɯɥɨɪɢɞɭ ɤɨɛɚɥɶɬɚ >@ ɤɚɪɨ ɬɢɧɨɢɞɨɜ± ɮɨɬɨɤɨɥɨɪɢɦɟɬɪɢɱɟɫɤɢɩɨɛɢɯɪɨɦɚɬɭɤɚɥɢɹ>@ɞɭɛɢɥɶɧɵɯɜɟɳɟɫɬɜɨɩɪɟɞɟɥɹɥɢɩɟɪɦɚɧ ɝɚɧɚɬɨɦɟɬɪɢɱɟɫɤɢɩɨɦɟɬɨɞɢɤɟȽɎ;,>@ɨɪɝɚɧɢɱɟɫɤɢɯɤɢɫɥɨɬ± ɦɟɬɨɞɨɦɧɟɣɬɪɚɥɢɡɚɰɢɢɩɨȽɎ;, ɫɩɪɢɦɟɧɟɧɢɟɦɦɟɬɢɥɟɧɨɜɨɝɨɫɢɧɟɝɨɜɨɞɨɪɚɫɬɜɨɪɢɦɵɯɩɨɥɢɫɚɯɚɪɢɞɨɜ± ɝɪɚɜɢɦɟɬɪɢɱɟɫɤɢ>]. 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Ɉɬɫɪɟɞɧɟɣɬɟɦɩɟɪɚɬɭɪɵ ɩɟɪɢɨɞ ɡɚɜɟɝɟɬɚɰɢɨɧɧɵɣɩɟɪɢɨɞ - 0,02 0,25 0,33 -0,22 -0,02 -0,20 -0,18 0,37 -0,34 0,37 -0,23 0,23 0,57 -0,15 ɂɡɬɚɛɥɢɰɵɜɢɞɧɨɱɬɨɡɚɜɢɫɢɦɨɫɬɶɮɥɚɜɨɧɨɢɞɨɜɨɬɡɧɚɱɟɧɢɹɨɫɚɞɤɨɜɡɚɜɟɝɟɬɚɰɢɨɧɧɵɣɩɟ ɪɢɨɞɨɛɪɚɬɧɚɹɚɨɬɬɟɦɩɟɪɚɬɭɪɵɩɪɹɦɚɹɜɨɬɥɢɱɢɟɨɬɞɚɧɧɵɯɩɨɫɨɞɟɪɠɚɧɢɸɚɫɤɨɪɛɢɧɨɜɨɣɤɢɫɥɨ ɬɵɇɚɞɨɨɬɦɟɬɢɬɶɱɬɨɤɨɪɪɟɥɹɰɢɹɫɨɞɟɪɠɚɧɢɹɮɥɚɜɨɧɨɢɞɨɜɢɚɧɬɨɰɢɚɧɨɜɨɬɨɫɚɞɤɨɜɯɨɬɹɢɨɬɪɢ ɰɚɬɟɥɶɧɚɹ ɧɨ ɩɪɚɤɬɢɱɟɫɤɢ ɧɟ ɡɧɚɱɢɬɟɥɶɧɚɹ ɉɪɨɫɥɟɠɢɜɚɟɬɫɹ ɨɛɳɚɹ ɬɟɧɞɟɧɰɢɹ ɩɨɜɵɲɟɧɢɹ ɫɨɞɟɪ ɠɚɧɢɹ ɮɥɚɜɨɧɨɢɞɨɜɢɞɪɭɝɢɯɮɟɧɨɥɶɧɵɯɫɨɟɞɢɧɟɧɢɣɡɚɢɫɤɥɸɱɟɧɢɟɦɚɧɬɨɰɢɚɧɨɜɨɬɬɟɦɩɟɪɚɬɭɪɵɢ ɫɧɢɠɟɧɢɟɢɯɫɨɞɟɪɠɚɧɢɹɨɬɜɥɚɝɨɨɛɟɫɩɟɱɟɧɧɨɫɬɢɯɨɬɹɨɧɨɩɪɨɹɜɥɹɟɬɫɹɜɦɟɧɶɲɟɣɫɬɟɩɟɧɢȼɨɛ ɳɟɦɦɨɠɧɨɫɞɟɥɚɬɶɜɵɜɨɞɱɬɨɧɟɦɚɥɨɜɚɠɧɭɸɪɨɥɶɢɝɪɚɟɬɬɟɦɩɟɪɚɬɭɪɚɜɨɡɞɭɯɚɢɜɥɚɝɨɨɛɟɫɩɟɱɟɧ ɧɨɫɬɶɪɚɫɬɟɧɢɣɜɫɢɧɬɟɡɟɢɨɛɦɟɧɟɮɥɚɜɨɧɨɢɞɧɵɯɪɚɫɬɟɧɢɣ ɇɚɞɨ ɨɬɦɟɬɢɬɶ ɱɬɨ ɞɨɫɬɚɬɨɱɧɚɹ ɜɥɚɝɨɨɛɟɫɩɟɱɟɧɧɨɫɬɶ ɜ ɟɫɬɟɫɬɜɟɧɧɵɯɭɫɥɨɜɢɹɯ ɫɬɢɦɭɥɢɪɭɟɬ ɫɢɧɬɟɡɚɫɤɨɪɛɢɧɨɜɨɣɤɢɫɥɨɬɵɢɩɨɥɢɫɚɯɚɪɢɞɨɜ Ʉɨɪɪɟɥɹɰɢɹ ɤɚɪɨɬɢɧɨɢɞɨɜ ɨɬ ɜɥɚɝɨɨɛɟɫɩɟɱɟɧɧɨɫɬɢ ɨɬɪɢɰɚɬɟɥɶɧɚɹ, ɯɨɬɹ ɢ ɩɪɨɹɜɥɹɟɬɫɹ ɜ ɦɟɧɶɲɟɣɫɬɟɩɟɧɢɚɨɬɬɟɦɩɟɪɚɬɭɪɵɩɨɥɨɠɢɬɟɥɶɧɚɹɌɟ, ɢɡɛɵɬɨɱɧɚɹɜɥɚɝɨɨɛɟɫɩɟɱɟɧɧɨɫɬɶɪɚɫɬɟɧɢɣ ɩɪɢɜɨɞɢɬɤɩɨɧɢɠɟɧɢɸɫɢɧɬɟɡɚɤɚɪɨɬɢɧɨɢɞɨɜɜɪɚɫɬɟɧɢɹɯ ɉɨɫɤɨɥɶɤɭɜɪɚɡɧɵɯɱɚɫɬɹɯɚɪɟɚɥɚɫɨɞɟɪɠɚɧɢɟɞɟɣɫɬɜɭɸɳɢɯɜɟɳɟɫɬɜɭɛɨɹɪɵɲɧɢɤɚɫɭɳɟɫɬ ɜɟɧɧɨɢɡɦɟɧɹɟɬɫɹɬɚɤɠɟɨɩɪɟɞɟɥɢɥɢɜɥɢɹɧɢɟɦɟɫɬɨɧɚɯɨɠɞɟɧɢɹɨɛɴɟɞɢɧɢɜɡɧɚɱɟɧɢɹɞɟɣɫɬɜɭɸɳɢɯ ɜɟɳɟɫɬɜ ɜ ɝɪɭɩɩɵ ɩɨ ɪɚɫɬɢɬɟɥɶɧɵɦ ɡɨɧɚɦ >@ ɢɥɢ ɛɨɥɟɟ ɨɛɨɛɳɟɧɧɨ ɩɨ ɞɜɭɦ ɪɚɣɨɧɚɦ ɫɟɜɟɪɨɡɚɩɚɞɧɵɣɩɪɟɞɝɨɪɧɵɣɢɰɟɧɬɪɚɥɶɧɵɣɩɪɟɞɝɨɪɧɵɣ [1]ȼɫɟɜɟɪɨ-ɡɚɩɚɞɧɵɣɩɪɟɞɝɨɪɧɵɣɪɚɣɨɧɜɨɲɥɢ ɨɤɪɟɫɬɧɨɫɬɢɫɫɍɱɤɟɧɬɗɪɩɟɥɢɇɨɜɨɥɚɤɫɤɨɟɝɆɚɯɚɱɤɚɥɵɜɰɟɧɬɪɚɥɶɧɵɣɩɪɟɞɝɨɪɧɵɣ± ɨɤɪɟɫɬ ɧɨɫɬɢɫɫȻɟɥɢɞɠɢȼɟɥɢɤɟɧɬɄɭɥɟɝɭ ɉɨɥɭɱɟɧɧɵɟɪɟɡɭɥɶɬɚɬɵɩɪɟɞɫɬɚɜɥɟɧɵɜɬɚɛɥ. 29 ɘɥɩɦɩɞɣɺ ɫɛɬɭɠɨɣɤ Ecology of plants əɞ ɋɩɬɬɣɣ: ɸɥɩɦɩɞɣɺ, ɫɛɢɝɣɭɣɠʌ 4, 2011 The South of Russia: ecology, development. ʋ4, 2011 Ɍɚɛɥɢɰɚ Ɂɚɜɢɫɢɦɨɫɬɶɫɨɞɟɪɠɚɧɢɹɛɢɨɥɨɝɢɱɟɫɤɢɚɤɬɢɜɧɵɯɜɟɳɟɫɬɜ ɜɩɥɨɞɚɯ Crataegus pentagyna Waldst et Kit ɨɬɦɟɫɬɨɧɚɯɨɠɞɟɧɢɹɜ-ɝ Ɋɟɫɭɪɫɧɵɟ ɪɚɣɨɧɵ ɋɟɜɟɪɨɡɚɩɚɞɧɵɣ ɩɪɟɞɝɨɪɧɵɣ ɐɟɧɬɪɚɥɶɧɵɣ ɩɪɟɞɝɨɪɧɵɣ ɋɟɜɟɪɨɡɚɩɚɞɧɵɣ ɩɪɟɞɝɨɪɧɵɣ ɐɟɧɬɪɚɥɶɧɵɣ ɩɪɟɞɝɨɪɧɵɣ ɋɟɜɟɪɨɡɚɩɚɞɧɵɣ ɩɪɟɞɝɨɪɧɵɣ ɐɟɧɬɪɚɥɶɧɵɣ ɩɪɟɞɝɨɪɧɵɣ Ƚɨɞ ɫɛɨɪɚ Ȼɢɨɥɨɝɢɱɟɫɤɢɚɤɬɢɜɧɵɟɜɟɳɟɫɬɜɚ ɚɫɤɨɪɛɢɧɨɜɚɹ ɮɥɚɜɨɧɨɢɞɵ ɚɧɬɨɰɢɚɧɵ ɤɢɫɥɨɬɚ ɋɪɟɞɧɢɟ ɋɢɥɚɢ ɋɪɟɞɧɢɟ ɋɢɥɚɢ ɋɪɟɞɧɢɟ ɋɢɥɚɢ ɞɚɧɧɵɟ ɞɨɫɬɨɜɟɪ ɞɚɧɧɵɟ ɞɨɫɬɨɜɟɪ ɞɚɧɧɵɟ ɞɨɫɬɨɜɟɪ ɩɨɝɪɭɩ ɜɢɹɧɢɹ ɩɨɝɪɭɩ ɜɢɹɧɢɹ ɩɨɝɪɭɩ ɜɢɹɧɢɹ ɩɟ ɮɚɤɬɨɪɚ ɩɟ ɮɚɤɬɨɪɚ ɩɟ ɮɚɤɬɨɪɚ 0,28 Ș2 = 0,83 0,63 Ș2 = 0,54 0,13 Ș2 = 0,33 F = 12,7 F = 3,2 F = 1,3 2001 0,17 0,26 2002 0,19 0,19 2003 0,17 Ș2 = 0,39 F = 1,9 Ș2 = 0,83 F =13,0 0,51 Ș2 = 0,89 F = 25,9 Ș2 = 0,75 F = 7,8 0,44 Ș2 = 0,37 F = 1,7 0,60 0,58 0,61 Ș2 = 0,42 F = 2,2 Ș2 = 0,81 F = 11,2 0,12 Ș2 = 0,92 F = 36,2 Ș2 = 0,79 F = 10,3 0,10 Ș2 = 0,66 F = 5,8 0,11 0,10 0,11 Ș2 = 0,15 F = 0,5 Ș2 = 0,52 F = 2,9 Ș2 = 0,22 F = 0,9 Ș2 = 0,23 F = 0,9 Ș2 = 0,05 F = 0,1 ɉɪɢɦɟɱɚɧɢɟF ± ɤɪɢɬɟɪɢɣɎɢɲɟɪɚȘ 2 ± ɫɢɥɚɜɥɢɹɧɢɹɮɚɤɬɨɪɚɍɪɨɜɟɧɶɞɨɫɬɨɜɟɪɧɨɫɬɢ± ɪ Ʉɚɤ ɜɢɞɧɨ ɢɡ ɬɚɛɥɢɰɵ ɦɟɫɬɨɧɚɯɨɠɞɟɧɢɟ ɨɤɚɡɵɜɚɟɬ ɞɨɫɬɚɬɨɱɧɨɟ ɫɢɥɶɧɨɟ ɜɥɢɹɧɢɟ ɧɚ ɫɨɞɟɪ ɠɚɧɢɟɮɥɚɜɨɧɨɢɞɨɜɢɚɫɤɨɪɛɢɧɨɜɨɣɤɢɫɥɨɬɵɜɩɥɨɞɚɯɛɨɹɪɵɲɧɢɤɚɩɹɬɢɩɟɫɬɢɱɧɨɝɨɦɚɤɫɢɦɚɥɶɧɵɟ ɫɨɞɟɪɠɚɧɢɹɤɨɬɨɪɵɯɨɬɦɟɱɟɧɵɭɛɨɹɪɵɲɧɢɤɚɩɹɬɢɩɟɫɬɢɱɧɨɝɨɩɪɨɢɡɪɚɫɬɚɸɳɟɝɨ ɜɫɟɜɟɪɨ-ɡɚɩɚɞɧɨɦ ɩɪɟɞɝɨɪɶɟɜɝɨɞɭ ȼɢɝɝɫɢɥɚɜɥɢɹɧɢɹɜɰɟɧɬɪɚɥɶɧɨɦɩɪɟɞɝɨɪɶɟɦɟɧɟɟ ɡɚɦɟɬɧɚɷɬɨɜɨɡɦɨɠɧɨ ɡɚɫɱɟɬɛɨɥɶɲɟɝɨɜɥɢɹɧɢɹɷɤɨɥɨɝɢɱɟɫɤɨɝɨɮɚɤɬɨɪɚ ɇɚ ɫɨɞɟɪɠɚɧɢɟ ɚɧɬɨɰɢɚɧɨɜ ɜ ɩɥɨɞɚɯ ɛɨɹɪɵɲɧɢɤɚ ɩɹɬɢɩɟɫɬɢɱɧɨɝɨ ɫɨɛɪɚɧɧɵɯ ɜ ɝɨɞɭ ɦɟɫɬɨɧɚɯɨɠɞɟɧɢɟɧɟɨɤɚɡɵɜɚɟɬɜɥɢɹɧɢɹɫɢɥɚɜɥɢɹɧɢɹ ± ɯɨɬɹɧɚɛɥɸɞɚɟɬɫɹɧɟɤɨɬɨɪɚɹɬɟɧɞɟɧ ɰɢɹ ɩɨɜɵɲɟɧɢɹ ɢɯɧɚɤɨɩɥɟɧɢɹ ɜ ɝɨɞɭ ɜ ɫɟɜɟɪɨ-ɡɚɩɚɞɧɨɦ ɩɪɟɞɝɨɪɶɟ ɋɢɥɶɧɨɟɜɥɢɹɧɢɟ (52 %) ɧɚɛɥɸɞɚɟɬɫɹɧɚɢɯɧɚɤɨɩɥɟɧɢɟɜɫɟɜɟɪɨ-ɡɚɩɚɞɧɨɦɩɪɟɞɝɨɪɶɟɜɝɨɞɭ Ⱦɥɹ ɜɵɹɜɥɟɧɢɹ ɜɥɢɹɧɢɹ ɷɤɨɥɨɝɢɱɟɫɤɢɯɮɚɤɬɨɪɨɜ ɬɚɤɢɯ ɤɚɤ ɬɟɦɩɟɪɚɬɭɪɚ ɢ ɤɨɥɢɱɟɫɬɜɨɨɫɚɞ ɤɨɜɫɪɚɜɧɢɥɢɨɛɪɚɡɰɵɩɥɨɞɨɜɛɨɹɪɵɲɧɢɤɚɫɨɛɪɚɧɧɵɯɫɬɪɟɯɦɟɫɬɩɪɨɢɡɪɚɫɬɚɧɢɹɨɤɪɟɫɬɧɨɫɬɢ ɫɫ Ƚɚɦɢɹɯɏɭɱɧɢɢɝɂɡɛɟɪɛɚɲɜɨɞɧɭɢɬɭɠɟɮɚɡɭɩɨɥɧɨɝɨɩɥɨɞɨɧɨɲɟɧɢɹɜɬɟɱɟɧɢɟɬɪɟɯɥɟɬ Ɍɚɛɥɢɰɚ ɋɨɞɟɪɠɚɧɢɟɛɢɨɥɨɝɢɱɟɫɤɢɚɤɬɢɜɧɵɯɜɟɳɟɫɬɜɜɩɥɨɞɚɯCrataegus pentagyna Waldst et Kit ɜɡɚɜɢɫɢɦɨɫɬɢɨɬɦɟɫɬɚɩɪɨɢɡɪɚɫɬɚɧɢɹɢɝɨɞɚɫɛɨɪɚ Ɇɟɫɬɨɫɛɨɪɚ ɫȽɚɦɢɹɯ ɫɏɭɱɧɢ ɝɂɡɛɟɪɛɚɲ ɫȽɚɦɢɹɯ ɫɏɭɱɧɢ ɝɂɡɛɟɪɛɚɲ ɫȽɚɦɢɹɯ ɫɏɭɱɧɢ ɝɂɡɛɟɪɛɚɲ 30 Ƚɨɞ ɫɛɨɪɚ 2006 2007 2008 ɋɭɦɦɚ ɨɫɚɞɤɨɜ ɡɚɜɟɝɟɬɚɰ ɩɟɪɢɨɞ 158,5 160,0 62,8 216,4 122,6 53,5 221,5 104,9 113,8 ɋɪɟɞɧɹɹ ɬɟɦɩɟɪɚɬɭɪɚ ɡɚɜɟɝɟɬɚɰɢɨɧ. ɩɟɪɢɨɞ 13,1 18,2 17,4 19,6 17,9 17,1 18,8 18,6 17,7 ɮɥɚɜɨ ɧɨɢɞɵ % Ʉɢɫɥɨɬɚ ɚɫɤɨɪɛ ɚɧɬɨɰɢɚ ɧɵ % 0,27 0,34 0,37 0,24 0,29 0,39 0,28 0,31 0,38 0,89 0,54 0,37 0,78 0,68 0,39 0,83 0,67 0,47 0,13 0,12 0,14 0,10 0,14 0,12 0,10 0,11 0,10 ɘɥɩɦɩɞɣɺ ɫɛɬɭɠɨɣɤ Ecology of plants əɞ ɋɩɬɬɣɣ: ɸɥɩɦɩɞɣɺ, ɫɛɢɝɣɭɣɠʌ 4, 2011 The South of Russia: ecology, development. ʋ4, 2011 ȼ ɰɟɥɨɦ ɟɫɥɢ ɫɪɚɜɧɢɜɚɬɶ ɫɪɟɞɧɸɸ ɬɟɦɩɟɪɚɬɭɪɭ ɡɚ ɜɟɝɟɬɚɰɢɨɧɧɵɣ ɩɟɪɢɨɞ ɛɨɹɪɵɲɧɢɤɚ ɢ ɫɭɦɦɭɨɫɚɞɤɨɜɜɵɩɚɜɲɭɸ ɡɚɷɬɨɬɩɟɪɢɨɞɜɦɟɫɬɚɯɫɛɨɪɚɦɨɠɧɨɫɤɚɡɚɬɶɱɬɨɜɫɟɨɬɦɟɱɟɧɧɵɟɝɨɞɵ ɛɵɥɢɛɥɚɝɨɩɪɢɹɬɧɵɦɢɞɥɹɛɨɹɪɵɲɧɢɤɚɏɨɬɹɧɚɢɦɟɧɶɲɟɟɤɨɥɢɱɟɫɬɜɨɨɫɚɞɤɨɜɜɵɩɚɜɲɟɟ ɡɚɩɟɪɢɨɞ ɜɟɝɟɬɚɰɢɢɜɨɬɦɟɱɟɧɧɵɟɝɨɞɵɜɨɤɪɟɫɬɧɨɫɬɹɯ ɫȽɚɦɢɹɯ ɧɚɛɥɸɞɚɟɬɫɹɜɝɨɞɭ, ɱɬɨɧɟɨɬɪɚɡɢɥɨɫɶ ɧɚɧɚɤɨɩɥɟɧɢɟȻȺȼ, ɩɨɫɤɨɥɶɤɭɧɚɛɥɸɞɚɟɦɵɣɩɟɪɢɨɞɛɵɥɞɨɜɨɥɶɧɨɬɚɤɢɩɪɨɯɥɚɞɧɵɦ Ɂɧɚɱɢɬɟɥɶɧɨɟ ɫɨɞɟɪɠɚɧɢɟ ɮɥɚɜɨɧɨɢɞɨɜ ɧɚɛɥɸɞɚɟɬɫɹ ɜ ɩɥɨɞɚɯ ɫ ɨɤɪɟɫɬɧɨɫɬɟɣ ɝ ɂɡɛɟɪɛɚɲ ɝɞɟɧɚɛɥɸɞɚɟɬɫɹɜɵɩɚɞɟɧɢɟɧɚɢɦɟɧɶɲɟɝɨɤɨɥɢɱɟɫɬɜɚɨɫɚɞɤɨɜɢɬɟɩɥɵɣɜɟɝɟɬɚɰɢɨɧɧɵɣɩɟɪɢɨɞɗɬɨ ɥɢɲɧɢɣɪɚɡɩɨɞɬɜɟɪɠɞɚɟɬɱɬɨɧɚɢɛɨɥɟɟɛɥɚɝɨɩɪɢɹɬɧɵɦɢɭɫɥɨɜɢɹɦɢɞɥɹɧɚɤɨɩɥɟɧɢɹɮɥɚɜɨɧɨɢɞɨɜ ɹɜɥɹɸɬɫɹ ɭɦɟɪɟɧɧɚɹ ɜɥɚɠɧɨɫɬɶ ɢ ɞɨɫɬɚɬɨɱɧɨ ɬɟɩɥɵɣ ɤɥɢɦɚɬ Ⱥ ɱɬɨ ɤɚɫɚɟɬɫɹ ɫɨɞɟɪɠɚɧɢɹ ɤɢɫɥɨɬɵ ɚɫɤɨɪɛɢɧɨɜɨɣɬɨ ɦɚɤɫɢɦɚɥɶɧɨɟɟɟɫɨɞɟɪɠɚɧɢɟɨɬɦɟɱɟɧɨɜɩɥɨɞɚɯɫɨɛɪɚɧɧɵɯɜɨɤɪɟɫɬɧɨɫɬɹɯ ɫȽɚ ɦɢɹɯ ɝɨɞɭɝɞɟɧɚɛɥɸɞɚɥɫɹɞɨɜɨɥɶɧɨɬɚɤɢɩɪɨɯɥɚɞɧɵɣɢɭɦɟɪɟɧɧɨɜɥɚɠɧɵɣɩɟɪɢɨɞɜɟɝɟɬɚɰɢɢ ɑɬɨ ɤɚɫɚɟɬɫɹ ɫɨɞɟɪɠɚɧɢɹ ɚɧɬɨɰɢɚɧɨɜ ɢɯ ɧɚɤɨɩɥɟɧɢɟ ɨɬ ɝɨɞɚ ɫɛɨɪɚ ɤɨɥɢɱɟɫɬɜɚ ɨɫɚɞɤɨɜ ɢ ɫɪɟɞɧɟɣ ɬɟɦɩɟɪɚɬɭɪɵ ɢɡɦɟɧɹɟɬɫɹ ɧɟɨɞɢɧɚɤɨɜɨ ɇɚɢɛɨɥɶɲɟɟ ɢɯ ɫɨɞɟɪɠɚɧɢɟ ɨɬɦɟɱɚɟɬɫɹ ɜ ɩɥɨɞɚɯ ɫɨɛɪɚɧɧɵɯɫɨɤɪɟɫɬɧɨɫɬɟɣ ɝɂɡɛɟɪɛɚɲɜɝɨɞɭɢɫɨɤɪɟɫɬɧɨɫɬɟɣ ɫɏɭɱɧɢɜɝɨɞɭɝɞɟɧɚ ɛɥɸɞɚɟɬɫɹɩɪɚɤɬɢɱɟɫɤɢɨɞɢɧɚɤɨɜɚɹɬɟɦɩɟɪɚɬɭɪɚɧɨɛɨɥɶɲɚɹɪɚɡɧɢɰɚɜɨɫɚɞɤɚɯɜɵɩɚɜɲɢɯɡɚɜɟɝɟ ɬɚɰɢɨɧɧɵɣɩɟɪɢɨɞ Ɋɟɡɭɥɶɬɚɬɵ ɬɚɛɥɢɰɵ ɩɨɤɚɡɵɜɚɸɬ ɱɬɨ ɫɨɞɟɪɠɚɧɢɟ ɛɢɨɥɨɝɢɱɟɫɤɢ ɚɤɬɢɜɧɵɯ ɜɟɳɟɫɬɜ ɨɬ ɝɨɞɚ ɫɛɨɪɚɤɨɥɢɱɟɫɬɜɚɨɫɚɞɤɨɜɢɫɪɟɞɧɟɣɬɟɦɩɟɪɚɬɭɪɵɡɚɜɟɝɟɬɚɰɢɨɧɧɵɣɩɟɪɢɨɞɛɨɹɪɵɲɧɢɤɚɢɡɦɟɧɹɟɬ ɫɹɧɟɨɞɢɧɚɤɨɜɨɢɧɟɡɧɚɱɢɬɟɥɶɧɨ ȼɣɜɦɣɩɞɫɛɯɣɲɠɬɥɣɤɬɪɣɬɩɥ ȺɤɚɟɜȻȺȺɬɚɟɜɁȼȽɚɞɠɢɟɜȻɋɢɞɪɎɢɡɢɱɟɫɤɚɹɝɟɨɝɪɚɮɢɹȾɚɝɟɫɬɚɧɚɆɒɤɨɥɚɫ 2. ȺɥɟɤɫɟɟɜȻȾɊɚɫɬɢɬɟɥɶɧɵɟɪɟɫɭɪɫɵȾɚɝɟɫɬɚɧɚɑɚɫɬɶɂɡɭɱɟɧɢɟɪɚɫɩɪɨɫɬɪɚɧɟɧɢɹɢɡɚɩɚɫɨɜɫɵɪɶɹ. ɍɱɟɛ ɧɨɟɩɨɫɨɛɢɟ Ɇɚɯɚɱɤɚɥɚ 3. ȺɬɥɚɫɊɟɫɩɭɛɥɢɤɢȾɚɝɟɫɬɚɧ. ɆɎɟɞɟɪɚɥɶɧɚɹɫɥɭɠɛɚɝɟɨɞɟɡɢɢɢɤɚɪɬɨɝɪɚɮɢɢɊɨɫɫɢɢ 4. ȻɟɪɝɅɋɈɫɧɨɜɵɤɥɢɦɚɬɨɥɨɝɢɢɂɡɞɆ-Ʌ. 5. ȽɨɧɱɚɪɨɜɚɌȺɗɧɰɢɤɥɨɩɟɞɢɹɥɟɤɚɪɫɬɜɟɧɧɵɯɪɚɫɬɟɧɢɣɅɟɱɟɧɢɟɬɪɚɜɚɦɢɆɂɡɞɚɬɟɥɶɫɤɢɣɞɨɦɆɋɉ 6. Ƚɨɫɭɞɚɪɫɬɜɟɧɧɚɹ Ɏɚɪɦɚɤɨɩɟɹ ɋɋɋɊ ȼɵɩ Ɉɛɳɢɟ ɦɟɬɨɞɵ ɚɧɚɥɢɡɚ Ʌɟɤɚɪɫɬɜɟɧɧɨɟ ɪɚɫɬɢɬɟɥɶɧɨɟ ɫɵɪɶɟ / ɆɁɋɋɋɊ-ɟɢɡɞɞɨɩɆɆɟɞɢɰɢɧɚ ɫ 7. Ƚɪɢɧɤɟɜɢɱ ɇɂɏɢɦɢɱɟɫɤɢɣɚɧɚɥɢɡɥɟɤɚɪɫɬɜɟɧɧɵɯɪɚɫɬɟɧɢɣɇɂȽɪɢɧɤɟɜɢɱɅɇɋɨɮɪɨɧɢɱ ɆɆɟɞɢ ɰɢɧɚɫ 8. ɄɚɡɚɤɨɜȺɅɆɟɬɨɞɵɢɫɫɥɟɞɨɜɚɧɢɹɩɪɢɪɨɞɧɵɯɮɥɚɜɨɧɨɢɞɨɜȺɅɄɚɡɚɤɨɜȺɅɒɢɧɤɚɪɟɧɤɨɆɟɬɨɞɢɱɟ ɫɤɢɟɪɟɤɨɦɟɧɞɚɰɢɢɞɥɹɫɬɭɞɟɧɬɨɜɧɚɭɱɧɨɝɨɫɬɭɞɟɧɱɟɫɤɨɝɨɨɛɳɟɫɬɜɚɉɹɬɢɝɨɪɫɤ 70 c. 9. Ʉɭɩɱɚɤ Ɍȼ Ʉɨɥɢɱɟɫɬɜɟɧɧɨɟ ɨɩɪɟɞɟɥɟɧɢɟ ɚɧɬɨɰɢɚɧɨɜ ɜ ɧɚɞɡɟɦɧɨɣ ɱɚɫɬɢ ɝɢɛɪɢɞɧɨɣ ɮɨɪɦɵ =HD 0D\V / ɌȼɄɭɩɱɚɤɅ$ɇɢɤɨɥɚɟɜɚɅɅɒɢɦɨɥɢɧɚ Ɋɚɫɬɢɬɟɥɶɧɵɟɪɟɫɭɪɫɵʋ ɋ-111. 10. ɄɨɱɤɚɪɨɜȼɂɎɚɪɦɚɤɨɝɧɨɫɬɢɱɟɫɤɨɟɢɡɭɱɟɧɢɟɧɟɤɨɬɨɪɵɯɩɪɟɞɫɬɚɜɢɬɟɥɟɣɫɟɦɟɣɫɬɜɚəɫɧɨɬɤɨɜɵɟȾɢɫ« ɤɚɧɞɮɚɪɦɚɰɧɚɭɤȼɂɄɨɱɤɚɪɨɜɄɭɪɫɤ 11. ɆɚɦɚɟɜɋȺɈɫɧɨɜɧɵɟɩɪɢɧɰɢɩɵɦɟɬɨɞɢɤɢɢɫɫɥɟɞɨɜɚɧɢɹɜɧɭɬɪɢɜɢɞɨɜɨɣɢɡɦɟɧɱɢɜɨɫɬɢɞɪɟɜɟɫɧɵɯɪɚɫɬɟ ɧɢɣ ɂɧɞɢɜɢɞɭɚɥɶɧɚɹɢɷɤɨɥɨɝɨ-ɝɟɨɝɪɚɮɢɱɟɫɤɚɹɢɡɦɟɧɱɢɜɨɫɬɶɪɚɫɬɟɧɢɣɋɜɟɪɞɥɨɜɫɤ C. 3-14. 12. ɉɪɢɫɬɭɩɚȿȺɋɨɜɟɪɲɟɧɫɬɜɨɜɚɧɢɟɚɧɚɥɢɡɚɢɬɟɯɧɨɥɨɝɢɢɢɡɝɨɬɨɜɥɟɧɢɹɧɚɫɬɨɟɜɲɢɩɨɜɧɢɤɚɤɪɚɩɢɜɵɢɪɹ ɛɢɧɵɨɛɵɤɧɨɜɟɧɧɨɣȿȺɉɪɢɫɬɭɩɚȾɆɉɨɩɨɜȾɆɆɭɯɚɦɟɞɠɚɧɨɜɚ ɎɚɪɦɚɰɢɹʋC. 23-25. Bibliography 1. Akaev B.A., Ataev Z.V. Gadzhiev B.S. Phizical geography of Dagestan. Moscow: Shcola, 1996. 386 p. 2. Alekseev B.D. Plant Resources of Dagestan. Part 2. Ɍhe study of distribution and stocks of raw materials. Textbook. Makhachkala, 1979. 3. Atlas of the Republic of Daghestan. Moscow: Federal Service of Geodesy and Cartography of Russia, 1999. 4. Berg, L.S. Foundations of Climatology. Publ. 2. Moscow-Leningrad, 1938. 5. Goncharova T.A. Encyclopedia of Medicinal Plants. Herbal therapy. Moscow: Publishing House of MSP, 1997. 6. State Pharmacopoeia of the USSR: Vol. 2. General methods of analysis. 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