трофические связи стрекоз (odonata)

ɘɥɩɦɩɞɣɺ ɡɣɝɩɭɨɶɰ
Ecology of animals
əɞ ɋɩɬɬɣɣ: ɸɥɩɦɩɞɣɺ, ɫɛɢɝɣɭɣɠ. ʌ1 2011
The South of Russia: ecology, development. ʋ
15. Bartenev A.N.Odonata outskirts of city Tiflis and other places Georgia // Bulletin of state museum of Georgia ± 1929ɛ. ± V. 5. ±
P. 65-86.
16. Bartenev A.N. About the spring and autumn fauna of dragonflies of Georgia and other places // Bulletin of state museum of
Georgia ± 1930ɚ. ± V. 6. ± P. 85-96.
17. Bartenev A.N. Notes about collectons of dragonflies North Caucasus of the hydrobiological station 1928 ɝ. in Kabarda and Balkaria // Works N. Caucasus of the hydrobiological station of Mountain rural-economic institute ± 1930ɛ. ± V. 3. ± pro. 1-3. ± P 85-86.
18. Bartenev A.N. Dragonflies of Western Caucasus // Labours 4 of All Union convention of zoologists, anatomists and histologists
Kiev 6-12 may 1930. ± Kiev-Harkov, 1931. ± P. 47-48.
19. Borisov S.N. Fauna and ecology of dragonflies of Tadjikistan Abstract of thesis of dissertation of candidate of biological
sciences ± Novosibirsk 1987. ± 23p.
20. Borisov S.N. About height distribution of dragonflies in Pamiro-Alai // Problems of ecology of mountain regions. ± Dushanbe,
1989. ± P.30-38.
21. Gorodkov K.B. Threedimensional climatic model of potenrial natural habitat and some her properties // Entomological commentator ± 1985. ± V. 64. ±Pro. 2. ± P.295-310.
22. Gorodkov K.B. Threedimensional climatic model of potential natural habitat and some her properties II // Entomological commentator - 1986. ± V. 65. ± Pro. 1. ± P.81-95.
23. Kozminov S.G., Ketenchiev H.A. Specific composition of dragonflies is in flat,foothill and mountain areas // Ecology and guard
of environment // Materials 4 of international conference - Ryazan,1998. ± P. 46-47.
24. Popova O.N. Dragonflies of family Sympetrum/ Diss. Of candidate of biological sciences Novosibirsk, 199. ± 261p.
25. Haritonov A.Y.Boreal fauna of dragonflies and ecology factor of geographical distribution Diss. doc. Biological sciences ± Novosibirsk, 1990. ± 536p.
26. Dumont H.J. Odonata of the Levant // Fauna Palaestina. Insecta V. ± Ierusalem, 1991. ± 304p.
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‹ 2011 ɄɟɬɟɧɱɢɟɜɏȺɌɢɯɨɧɨɜɚȺȼ
Ʉɚɛɚɪɞɢɧɨ-Ȼɚɥɤɚɪɫɤɢɣ ɝɨɫɭɞɚɪɫɬɜɟɧɧɵɣ ɭɧɢɜɟɪɫɢɬɟɬ
ȼ ɫɬɚɬɶɟ ɪɚɫɫɦɚɬɪɢɜɚɸɬɫɹ ɬɪɨɮɢɱɟɫɤɢɟ ɫɜɹɡɢ ɫɬɪɟɤɨɡ ɜ ɛɢɨɰɟɧɨɡɚɯ ɐɟɧɬɪɚɥɶɧɨɝɨ Ʉɚɜɤɚɡɚ Ⱥɧɚɥɢɡ ɢɡɭɱɚɟɦɨɝɨ
ɹɜɥɟɧɢɹɩɨɡɜɨɥɢɥɜɵɹɜɢɬɶɫɩɟɤɬɪɞɚɧɧɵɯɫɜɹɡɟɣɢɦɚɝɨɢɥɢɱɢɧɨɤɩɪɟɞɫɬɚɜɢɬɟɥɟɣɨɬɪɹɞɚ2GRQDWDɜɫɨɨɛɳɟɫɬɜɚɯ
ɢɫɫɥɟɞɭɟɦɨɝɨɪɚɣɨɧɚɋɬɪɭɤɬɭɪɚ ɬɪɨɮɢɱɟɫɤɢɯ ɫɜɹɡɟɣ ɩɪɟɞɫɬɚɜɥɟɧɚ ɜ ɜɢɞɟ ɞɢɚɝɪɚɦɦ.
The trophic links of dragonflies in biocenoses of the Central Caucasus are examined in this article. The analysis of this
pheQRPHQRQDOORZHGWRUHYHDOWKHVSHFWUXPRIWKHJLYHQFRQQHFWLRQRILPDJRDQGODUYD¶VPHPEHUVRIWKHRUGHU2GRQDWD
in the communities of the study area. The structure of the trophic links is represented in the form of diagrams.
Ʉɥɸɱɟɜɵɟɫɥɨɜɚ ɫɬɪɟɤɨɡɚɛɢɨɰɟɧɨɡɫɨɨɛɳɟɫɬɜɨɠɢɜɵɯɨɪɝɚɧɢɡɦɨɜɬɪɨɮɢɱɟɫɤɚɹɫɜɹɡɶ
Key words: dragonfly, biocenosis, community of living organisms, trophic link.
ȼɛɢɨɰɟɧɨɡɟɬɪɨɮɢɱɟɫɤɢɟɢɬɨɩɢɱɟɫɤɢɟɫɜɹɡɢɢɝɪɚɸɬɪɟɲɚɸɳɭɸɪɨɥɶɢɹɜɥɹɸɬɫɹɨɫɧɨ
ɜɨɣɟɝɨɫɭɳɟɫɬɜɨɜɚɧɢɹɗɬɢɨɬɧɨɲɟɧɢɹɭɞɟɪɠɢɜɚɸɬɨɪɝɚɧɢɡɦɵɪɚɡɧɵɯɜɢɞɨɜɨɛɴɟɞɢɧɹɹɢɯɜ
ɞɨɫɬɚɬɨɱɧɨɫɬɚɛɢɥɶɧɵɟɫɨɨɛɳɟɫɬɜɚɌɪɨɮɢɱɟɫɤɢɟɫɜɹɡɢɧɚɛɥɸɞɚɸɬɫɹɤɨɝɞɚɨɞɢɧɜɢɞɩɢɬɚɟɬɫɹ
ɩɪɟɞɫɬɚɜɢɬɟɥɹɦɢɞɪɭɝɨɝɨɥɢɛɨɢɯɦɟɪɬɜɵɦɢɨɫɬɚɬɤɚɦɢɥɢɛɨɩɪɨɞɭɤɬɚɦɢɠɢɡɧɟɞɟɹɬɟɥɶɧɨɫɬɢ
ɗɬɨ ɫɥɭɱɚɢ ɩɪɹɦɨɣ ɬɪɨɮɢɱɟɫɤɨɣ ɫɜɹɡɢ ɉɪɢ ɤɨɧɤɭɪɟɧɰɢɢ ɞɜɭɯ ɜɢɞɨɜ ɢɡ-ɡɚ ɨɛɴɟɤɬɨɜ ɩɢɬɚɧɢɹ
ɦɟɠɞɭɧɢɦɢɜɨɡɧɢɤɚɟɬɤɨɫɜɟɧɧɚɹɬɪɨɮɢɱɟɫɤɚɹɫɜɹɡɶɜɫɥɟɞɫɬɜɢɟɬɨɝɨɱɬɨɞɟɹɬɟɥɶɧɨɫɬɶɨɞɧɨɝɨ
ɜɢɞɚɨɬɪɚɠɚɟɬɫɹɧɚɫɧɚɛɠɟɧɢɢɤɨɪɦɨɦɞɪɭɝɨɝɨ
ɋɨɫɬɚɜ ɩɢɳɢ ɩɪɟɞɫɬɚɜɢɬɟɥɟɣ ɨɬɪɹɞɚ OGRQDWD ɢɡɭɱɟɧ ɧɟɞɨɫɬɚɬɨɱɧɨ Ɉɞɧɚɤɨ ɢɡɜɟɫɬɧɨ
ɱɬɨɷɬɢɯɢɳɧɢɤɢɢɫɩɨɥɶɡɭɸɬɜɤɚɱɟɫɬɜɟɩɢɳɢɥɸɛɨɟɧɚɫɟɤɨɦɨɟɫɤɨɬɨɪɵɦɦɨɝɭɬɫɩɪɚɜɢɬɶɫɹ
ɋɬɪɟɤɨɡɵɤɚɤɛɢɨɥɨɝɢɱɟɫɤɢɟɨɛɴɟɤɬɵɜɯɨɞɹɬɜɫɨɫɬɚɜɛɢɨɰɟɧɨɡɨɜɜɤɨɬɨɪɵɯɞɟɣɫɬɜɭɸɬɩɟɪɟ
ɱɢɫɥɟɧɧɵɟɜɡɚɢɦɨɨɬɧɨɲɟɧɢɹɜɧɨɫɹɫɭɳɟɫɬɜɟɧɧɵɣɜɤɥɚɞɜɢɯɫɬɪɭɤɬɭɪɭ
ȼɪɚɦɤɚɯɤɨɦɩɥɟɤɫɧɨɝɨɢɡɭɱɟɧɢɹɷɤɨɥɨɝɢɢɢɛɢɨɥɨɝɢɢɩɪɟɞɫɬɚɜɢɬɟɥɟɣɨɬɪɹɞɚ Odonata
ɰɟɥɶɸ ɧɚɲɟɝɨ ɢɫɫɥɟɞɨɜɚɧɢɹ ɫɬɚɥɨ ɜɵɹɜɥɟɧɢɟ ɬɪɨɮɢɱɟɫɤɢɯ ɫɜɹɡɟɣ ɫɬɪɟɤɨɡ ɜ ɛɢɨɰɟɧɨɡɚɯ ɐɟɧ
ɬɪɚɥɶɧɨɝɨɄɚɜɤɚɡɚ
Ɉɛɴɟɤɬɚɦɢɩɨɫɥɭɠɢɥɢɫɨɨɛɳɟɫɬɜɚɠɢɜɵɯɨɪɝɚɧɢɡɦɨɜɜɫɨɫɬɚɜɤɨɬɨɪɵɯɜɤɚɱɟɫɬɜɟɨɞ
ɧɨɝɨɢɡɫɭɳɟɫɬɜɟɧɧɵɯɡɜɟɧɶɟɜɜɯɨɞɹɬɫɬɪɟɤɨɡɵ
ɂɫɫɥɟɞɨɜɚɧɢɹɩɪɨɜɨɞɢɥɢɫɶɫɩɨɝɝȼɟɥɫɹɚɧɚɥɢɡɫɩɟɤɬɪɚɬɪɨɮɢɱɟɫɤɢɯɫɜɹɡɟɣ
ɢɦɚɝɨ ɢ ɥɢɱɢɧɨɤ ɩɪɟɞɫɬɚɜɢɬɟɥɟɣ ɷɬɨɝɨ ɨɬɪɹɞɚ ɜ ɩɪɟɞɟɥɚɯ ɪɚɡɥɢɱɧɵɯ ɜɵɫɨɬɧɵɯ ɩɨɹɫɨɜ ɐɟɧ
ɬɪɚɥɶɧɨɝɨɄɚɜɤɚɡɚɆɚɬɟɪɢɚɥɨɦɞɥɹɞɚɧɧɨɣɪɚɛɨɬɵɩɨɫɥɭɠɢɥɢɷɤɫɩɟɞɢɰɢɨɧɧɵɟɢɫɫɥɟɞɨɜɚɧɢɹ
88
ɘɥɩɦɩɞɣɺ ɡɣɝɩɭɨɶɰ
Ecology of animals
əɞ ɋɩɬɬɣɣ: ɸɥɩɦɩɞɣɺ, ɫɛɢɝɣɭɣɠ. ʌ1 2011
The South of Russia: ecology, development. ʋ
ɩɪɨɜɨɞɢɜɲɢɟɫɹ ɚɜɬɨɪɚɦɢ ɧɚ ɬɟɪɪɢɬɨɪɢɢ ɪɚɫɫɦɚɬɪɢɜɚɟɦɨɝɨ ɪɚɣɨɧɚ Ɍɪɨɮɢɱɟɫɤɢɟ ɫɜɹɡɢ ɭɫɬɚ
ɧɚɜɥɢɜɚɥɢɫɶɜɬɢɩɨɜɵɯɛɢɨɬɨɩɚɯɩɭɬɟɦɩɪɹɦɨɝɨɧɚɛɥɸɞɟɧɢɹɡɚɫɬɪɟɤɨɡɚɦɢɈɬɥɨɜɷɤɡɟɦɩɥɹɪɨɜ
ɨɫɭɳɟɫɬɜɥɹɥɫɹɜɨɡɞɭɲɧɵɦɫɚɱɤɨɦɢɦɚɝɨɢɛɢɨɰɟɧɨɦɟɬɪɨɦɩɪɨɫɬɟɣɲɟɣɤɨɧɫɬɪɭɤɰɢɢ± ɜɜɢɞɟ
ɦɟɬɚɥɥɢɱɟɫɤɨɣɬɪɭɛɵɤɜɚɞɪɚɬɧɨɝɨɫɟɱɟɧɢɹɫɨɫɬɨɪɨɧɨɣɫɦɢɜɵɫɨɬɨɣɨɬɞɨɫɦɥɢ
ɱɢɧɤɢɍɱɟɬɩɪɨɜɨɞɢɥɫɹɱɟɪɟɡɤɚɠɞɵɟɞɧɟɣɉɪɢɨɩɪɟɞɟɥɟɧɢɢɢɫɩɨɥɶɡɨɜɚɥɢɨɩɪɟɞɟɥɢɬɟɥɢ
ɫɬɪɟɤɨɡȻɎȻɟɥɵɲɟɜɚȻɎȻɟɥɵɲɟɜɚȺɘɏɚɪɢɬɨɧɨɜɚɏȺɄɟɬɟɧɱɢɟɜɚȺɘɏɚɪɢɬɨɧɨ
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ɋɩɟɤɬɪɬɪɨɮɢɱɟɫɤɢɯɫɜɹɡɟɣɫɬɪɟɤɨɡɤɚɤɢɞɪɭɝɢɯɨɪɝɚɧɢɡɦɨɜ ɜɛɢɨɰɟɧɨɡɚɯɐɟɧɬɪɚɥɶ
ɧɨɝɨɄɚɜɤɚɡɚ ɲɢɪɨɤɢɜɟɫɶɦɚɦɧɨɝɨɨɛɪɚɡɟɧ ɪɢɫɈɧ ɨɩɪɟɞɟɥɹɟɬɫɹɯɢɳɧɢɱɟɫɤɨɣɩɪɢɪɨɞɨɣ
ɢɯɩɨɜɟɞɟɧɢɹɨɛɴɟɦɨɦɜɢɞɨɜɨɝɨɪɚɡɧɨɨɛɪɚɡɢɹɚɬɚɤɠɟɦɟɫɬɨɦɨɛɢɬɚɧɢɹɥɢɱɢɧɨɱɧɨɣɮɚɡɵɪɚɡ
ɜɢɬɢɹɄɚɤɜɡɪɨɫɥɵɟɩɪɟɞɫɬɚɜɢɬɟɥɢɨɬɪɹɞɚɬɚɤɢɥɢɱɢɧɤɢɹɜɥɹɸɬɫɹɯɢɳɧɢɤɚɦɢ
ɉɢɳɚ
50
ɦɟɥɤɢɟɪɚɤɨɨɨɛɪɚɡɧɵɟ
45
ɦɟɥɤɢɟɧɚɫɟɤɨɦɵɟɢɢɯɥɢɱɢɧɤɢ
40
ɦɨɥɨɞɶɪɵɛ
ɞɪɭɝɢɟɨɛɴɟɤɬɵ
35
ȼɪɚɝɢ
30
ɦɨɥɨɞɶɪɵɛ
25
ɨɤɨɥɨɜɨɞɧɵɟɩɬɢɰɵ
20
ɦɥɟɤɨɩɢɬɚɸɳɢɟ
ɪɟɩɬɢɥɢɢ
15
ɉɚɪɚɡɢɬɵ
10
ɦɟɬɚɰɟɪɤɚɪɢɢɬɪɟɦɚɬɨɞ ɩɪɨɦɟɠɭɬɨɱɧɵɟɯɨɡɹɟɜɚ
ɜɨɞɧɵɟɤɥɟɳɢɢɢɯɥɢɱɢɧɤɢ
5
ɝɪɟɝɚɪɢɧɵ
0
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ȼɪɚɝɢ
ɉɚɪɚɡɢɬɵ
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ɧɟɦɚɬɨɞɵ
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40
̡̨̨̨̡̨̛̬̭̭̱̺̖̦̭̖̥̼̖̏̌
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35
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̸̡̨̡̛̛̥̖̣̖̍̌̍
30
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25
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20
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15
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10
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5
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;̸̵̨̨̨̪̬̥̖̙̱̯̦̼̖̖́̏̌̚Ϳ
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89
ɘɥɩɦɩɞɣɺ ɡɣɝɩɭɨɶɰ
Ecology of animals
əɞ ɋɩɬɬɣɣ: ɸɥɩɦɩɞɣɺ, ɫɛɢɝɣɭɣɠ. ʌ1 2011
The South of Russia: ecology, development. ʋ
ɂɦɚɝɨɫɬɪɟɤɨɡ ɜɟɞɭɬɞɧɟɜɧɨɣɨɛɪɚɡɠɢɡɧɢɥɟɬɚɸɬ ɩɪɟɢɦɭɳɟɫɬɜɟɧɧɨɜɹɫɧɭɸɩɨɝɨɞɭ
ɩɪɨɜɨɞɹ ɛɨɥɶɲɭɸ ɱɚɫɬɶ ɜɪɟɦɟɧɢ ɜ ɩɨɢɫɤɚɯ ɢ ɞɨɛɵɱɟ ɩɢɳɢ Ɂɚɮɢɤɫɢɪɨɜɚɧ ɮɚɤɬ ɩɨɟɞɚɧɢɹ
ɜɡɪɨɫɥɵɦɢ ɫɬɪɟɤɨɡɚɦɢ ɤɪɭɩɧɵɯ ɧɚɫɟɤɨɦɵɯ ɫɥɟɩɧɟɣ ɛɚɛɨɱɟɤ ɪɭɱɟɣɧɢɤɨɜ ɉɢɳɟɣ ɥɢɱɢɧɨɤ
ɫɥɭɠɚɬɦɟɥɤɢɟɪɚɤɨɨɛɪɚɡɧɵɟɞɚɮɧɢɢɚɦɮɢɩɨɞɵɰɢɤɥɨɩɵɦɟɥɤɢɟɧɚɫɟɤɨɦɵɟɢɢɯɥɢɱɢɧɤɢ
ɝɨɥɨɜɚɫɬɢɤɢɢɦɨɥɨɞɶɪɵɛ
ɏɚɪɚɤɬɟɪɧɨɣ ɨɫɨɛɟɧɧɨɫɬɶɸ ɤɪɭɩɧɵɯ ɜɢɞɨɜ ɫɬɪɟɤɨɡ Anax imperator, Anax parthenope,
Aeshna affinis, Aeshna cyanea, Aeshna mixta, Orthetrum brunneum, Orthetrum cancellatum, Crocothemis erythrae, Sympetrum vulgatum, Sympetrum pedemontanum, Sympetrum fonscolombei, Sympetrum meridionale, Libellula depressa, Libellula quadrimaculata, Lindenia tetraphilla) ɹɜɥɹɟɬɫɹ ɫɨ
ɜɟɪɲɟɧɢɟɠɢɪɨɜɨɱɧɵɯɩɨɥɟɬɨɜɍɬɚɤɢɯɩɪɟɞɫɬɚɜɢɬɟɥɟɣɧɚɛɥɸɞɚɟɬɫɹɯɨɪɨɲɨɜɵɪɚɠɟɧɧɚɹɬɟɪ
ɪɢɬɨɪɢɚɥɶɧɨɫɬɶ ɡɚɤɥɸɱɚɸɳɚɹɫɹ ɜ ɧɚɥɢɱɢɢ ɢɧɞɢɜɢɞɭɚɥɶɧɵɯ ɤɨɪɦɨɜɵɯ ɭɱɚɫɬɤɨɜ Ɇɟɥɤɢɟ ɠɟ
ɜɢɞɵ ɨɯɨɬɹɬɫɹ ɢɡ ɡɚɫɚɞɵ, ɱɬɨ ɨɫɨɛɟɧɧɨ ɹɪɤɨ ɩɪɨɹɜɥɹɟɬɫɹɜɱɚɫɬɧɨɫɬɢɭ ɩɪɟɞɫɬɚɜɢɬɟɥɟɣLestses
viridis, Lestes sponsa, Sympecma fusca, Coenagrion pulchellum Erythromma viridulum, Ischnura
elegans, Enallagma cyathigerum. ȼɫɥɭɱɚɟɠɟɟɫɥɢɷɬɨɧɟɩɪɢɧɨɫɢɬɪɟɡɭɥɶɬɚɬɨɜɨɧɢɩɟɪɟɥɟɬɚɸɬ
ɧɚɞɪɭɝɨɟɦɟɫɬɨ. ɍɜɢɞɨɜɫɩɨɞɨɛɧɵɦɢɨɫɨɛɟɧɧɨɫɬɹɦɢɬɪɨɮɢɱɟɫɤɨɣ ɚɤɬɢɜɧɨɫɬɢɢɧɞɢɜɢɞɭɚɥɶ
ɧɵɯ ɤɨɪɦɨɜɵɯ ɭɱɚɫɬɤɨɜɧɚɦ ɜɵɹɜɢɬɶ ɧɟ ɭɞɚɥɨɫɶ Ⱦɚɧɧɨɟ ɨɛɫɬɨɹɬɟɥɶɫɬɜɨ ɨɛɴɹɫɧɹɟɬɫɹ ɫɤɭɱɟɧ
ɧɨɫɬɶɸ ɨɫɨɛɟɣ ɧɚ ɬɟɪɪɢɬɨɪɢɢ ɤɨɪɦɥɟɧɢɹ ɜɛɥɢɡɢ ɜɨɞɨɟɦɨɜ ɢɥɢ ɠɟ ɟɞɢɧɢɱɧɨɫɬɶɸ ɜɫɬɪɟɱ ɧɚ
ɡɧɚɱɢɬɟɥɶɧɨɦɭɞɚɥɟɧɢɢɨɬɧɢɯ
ɇɚɛɥɸɞɟɧɢɹɩɨɤɚɡɚɥɢɧɚɥɢɱɢɟɹɜɥɟɧɢɹɦɚɫɫɨɜɨɝɨɢɫɬɪɟɛɥɟɧɢɹɫɬɪɟɤɨɡɚɦɢɦɟɥɤɢɯɤɪɨ
ɜɨɫɨɫɭɳɢɯɧɚɫɟɤɨɦɵɯɤɨɬɨɪɨɟɜɛɨɥɶɲɟɣɫɬɟɩɟɧɢɩɪɨɹɜɥɹɟɬɫɹɭAeshna affinis, Aeshna cyanea,
Aeshna grandis, Aeshna mixta, Anaciaeshna isosceles, Anax imperator, Anax parthenopeɌɨɟɫɬɶɭ
ɤɪɭɩɧɵɯɩɪɟɞɫɬɚɜɢɬɟɥɟɣɩɨɞɨɬɪɹɞɚ$QL]RSWHUD ɊɚɡɧɨɤɪɵɥɵɟɫɬɪɟɤɨɡɵAnizoptera) ɱɚɳɟɥɨɜɹɬ
ɧɚɫɟɤɨɦɵɯɜɜɨɡɞɭɯɟɚɪɚɜɧɨɤɪɵɥɵɟZygopteraɫɨɛɢɪɚɸɬɫɥɢɫɬɶɟɜɫɬɟɛɥɟɣɬɪɚɜɢɤɭɫɬɚɪɧɢɤɨɜ
ɇɚɦɢɨɬɦɟɱɟɧɩɪɨɰɟɫɫɩɨɟɞɚɧɢɹɛɨɥɟɟɦɟɥɤɢɯɫɬɪɟɤɨɡɤɪɭɩɧɵɦɢɩɪɟɞɫɬɚɜɢɬɟɥɹɦɢɞɪɭ
ɝɢɯ ɜɢɞɨɜ ɜ ɱɚɫɬɧɨɫɬɢ ɨɫɨɛɟɣ ɜɢɞɚ Enallagma cyathigerum ɩɪɟɜɨɫɯɨɞɹɳɢɦɢ ɢɯ ɩɨ ɪɚɡɦɟɪɚɦ
ɨɫɨɛɹɦɢɜɢɞɨɜAnax imperator ɢAnax parthenopeɩɪɢɛɨɥɶɲɨɣɩɥɨɬɧɨɫɬɢɫɨɨɛɳɟɫɬɜɫɩɪɟɨɛ
ɥɚɞɚɧɢɟɦɛɨɥɟɟɤɪɭɩɧɵɯɮɨɪɦɉɨɞɨɛɧɵɟɮɚɤɬɵɩɨɞɬɜɟɪɠɞɚɸɬɫɹɢɥɢɬɟɪɚɬɭɪɧɵɦɢɞɚɧɧɵɦɢ
ɇɚɩɪɢɦɟɪȺȺɋɥɭɜɤɨ, ȿɋɑɭɣɤɨɜɡɚɮɢɤɫɢɪɨɜɚɥɢɩɪɨɰɟɫɫɩɨɟɞɚɧɢɹ ɩɪɟɞɫɬɚɜɢɬɟɥɹɦɢ ɜɢɞɚ
Orthetrum cancellatum ɦɨɥɨɞɵɯ ɨɫɨɛɟɣ Crocothemis erythraea, ɚ ɩɪɟɞɫɬɚɜɢɬɟɥɹɦɢ Crocothemis
erythraea ɨɫɨɛɟɣ Sympetrum sanguineum ɤɨɬɨɪɵɟ ɭɫɬɭɩɚɸɬ ɢɦ ɜ ɪɚɡɦɟɪɚɯ [9]. ȼ ɬɨ ɠɟ ɜɪɟɦɹ
ɧɟɤɨɬɨɪɵɟ ɭɱɟɧɵɟ ɨɬɪɢɰɚɸɬ ɫɭɳɟɫɬɜɨɜɚɧɢɟ ɤɚɧɧɢɛɚɥɢɡɦɚ ɭ ɫɬɪɟɤɨɡ ɬɚɤɨɜɚ ɩɨɡɢɰɢɹ S. H.
Chowdhury, P. S. Corbet [10@Ⱦɪɭɝɢɟɢɫɫɥɟɞɨɜɚɬɟɥɢ ɨɬɦɟɱɚɸɬɱɬɨɧɟɫɬɚɥɤɢɜɚɥɢɫɶɫɩɨɞɨɛɧɵɦ
ɹɜɥɟɧɢɟɦ[11@Ɉɞɧɚɤɨɨɧɨɜɢɞɢɦɨɹɜɥɹɟɬɫɹɨɞɧɨɣɢɡɮɨɪɦɫɞɟɪɠɢɜɚɧɢɹɤɨɧɤɭɪɟɧɰɢɢɡɚɩɢ
ɳɟɜɵɟɪɟɫɭɪɫɵɜɦɟɫɬɚɯɦɚɫɫɨɜɨɝɨɫɤɨɩɥɟɧɢɹɫɬɪɟɤɨɡɫɩɪɟɨɛɥɚɞɚɧɢɟɦɛɨɥɟɟɤɪɭɩɧɵɯɩɪɟɞ
ɫɬɚɜɢɬɟɥɟɣɨɬɪɹɞɚɚɬɚɤɠɟɩɪɢɨɛɨɫɬɪɟɧɢɢɛɨɪɶɛɵɡɚɨɫɜɨɟɧɢɟɦɟɫɬɨɯɨɬɵɢɭɤɪɵɬɢɣɨɬɯɢɳ
ɧɢɤɨɜɩɪɢɪɟɡɤɨɦɭɜɟɥɢɱɟɧɢɢɱɢɫɥɟɧɧɨɫɬɢɜɢɞɨɜɜɯɨɞɹɳɢɯɜɫɨɫɬɚɜɨɞɨɧɚɬɨɤɨɦɩɥɟɤɫɚɤɚɤɨɣɥɢɛɨɬɟɪɪɢɬɨɪɢɢ
Ʌɢɱɢɤɢɫɬɪɟɤɨɡ- ɜɚɠɧɚɹɫɨɫɬɚɜɥɹɸɳɚɹ ɤɨɪɦɨɜɨɣɛɚɡɵɜɨɞɨɩɥɚɜɚɸɳɢɯɢɨɤɨɥɨɜɨɞɧɵɯ
ɩɬɢɰ ɪɵɛ ɢ ɢɯ ɦɨɥɨɞɢ ȼ ɬɨ ɠɟ ɜɪɟɦɹ ɫɥɟɞɭɟɬ ɩɨɞɱɟɪɤɧɭɬɶ ɱɬɨ ɧɚɪɹɞɭ ɫ ɷɬɢɦ ɨɧɢ ɩɨɟɞɚɸɬ
ɦɨɥɨɞɶɪɵɛɚɬɚɤɠɟɩɢɬɚɹɫɶɦɟɥɤɢɦɢɪɚɤɨɨɛɪɚɡɧɵɦɢɦɟɥɤɢɦɢɧɚɫɟɤɨɦɵɦɢɢɢɯɥɢɱɢɧɤɚɦɢ
ɤɨɧɤɭɪɢɪɭɸɬɫɧɟɣɡɚɢɫɩɨɥɶɡɨɜɚɧɢɟɩɢɳɟɜɵɯɪɟɫɭɪɫɨɜ.
ɉɪɟɞɫɬɚɜɢɬɟɥɢɨɬɪɹɞɚ2GRQDWDɫɥɭɠɚɬɦɨɳɧɵɦɮɚɤɬɨɪɨɦɪɟɝɭɥɢɪɭɸɳɢɦɱɢɫɥɟɧɧɨɫɬɶ
ɥɢɱɢɧɨɤ ɦɨɲɟɤ ɤɨɦɚɪɨɜ ɢ ɦɧɨɝɢɯ ɦɟɥɤɢɯ ɧɚɫɟɤɨɦɵɯ ɜɟɞɭɳɢɯ ɜɨɞɧɵɣ ɢ ɩɨɥɭɜɨɞɧɵɣ ɨɛɪɚɡ
ɠɢɡɧɢ ɢ ɞɪɭɝɢɯ ɨɛɢɬɚɬɟɥɟɣ ɜɨɞɨɟɦɨɜ ɬɟɦ ɫɚɦɵɦ ɨɛɟɫɩɟɱɢɜɚɹ ɩɨɞɞɟɪɠɚɧɢɟ ɷɤɨɥɨɝɢɱɟɫɤɨɝɨ
ɪɚɜɧɨɜɟɫɢɹ ɜ ɛɢɨɰɟɧɨɡɚɯ ɍɱɢɬɵɜɚɹ ɩɥɨɬɧɨɫɬɶ ɡɚɫɟɥɟɧɢɹ ɫɬɪɟɤɨɡɚɦɢ ɜɨɞɨɟɦɨɜ ɐɟɧɬɪɚɥɶɧɨɝɨ
Ʉɚɜɤɚɡɚɦɨɠɧɨɡɚɤɥɸɱɢɬɶɱɬɨɷɬɢɧɚɫɟɤɨɦɵɟɫɨɫɬɚɜɥɹɸɬɡɧɚɱɢɬɟɥɶɧɭɸɱɚɫɬɶɛɢɨɥɨɝɢɱɟɫɤɢɯ
ɪɟɫɭɪɫɨɜɪɟɝɢɨɧɚ
ɋɬɪɟɤɨɡɵɢɦɟɸɬɞɨɫɬɚɬɨɱɧɨɦɧɨɝɨɱɢɫɥɟɧɧɵɯɟɫɬɟɫɬɜɟɧɧɵɯɜɪɚɝɨɜ ɋɪɟɞɢɧɢɯɨɫɨɛɨɝɨ
ɜɧɢɦɚɧɢɹ ɡɚɫɥɭɠɢɜɚɸɬ ɨɪɝɚɧɢɡɦɵ ɨɬɧɨɫɹɳɢɟɫɹ ɤ ɧɟɫɤɨɥɶɤɢɦ ɫɢɫɬɟɦɚɬɢɱɟɫɤɢɦ ɝɪɭɩɩɚɦ ɧɟ
ɦɚɬɨɞɵ ɝɨɪɝɨɞɟɪɢɞɵ ɝɪɟɝɚɪɢɧɵ ɥɢɱɢɧɤɢ ɰɟɫɬɨɞ ɥɢɱɢɧɤɢ ɜɨɞɧɵɯ ɤɥɟɳɟɣ ɢ ɤɥɟɳɟɣɤɪɚɫɧɨɬɟɥɨɤ Ʉɪɨɦɟ ɬɨɝɨ ɢɦɚɝɨ ɢ ɥɢɱɢɧɤɢ ɩɪɟɞɫɬɚɜɢɬɟɥɟɣ ɨɬɪɹɞɚ 2donata - ɩɪɨɦɟɠɭɬɨɱɧɵɟ
ɯɨɡɹɟɜɚ ɧɟɤɨɬɨɪɵɯ ɜɢɞɨɜ ɬɪɟɦɚɬɨɞ. ɋɜɨɛɨɞɧɵɟ ɦɟɬɚɰɟɪɤɚɪɢɢ ɷɬɢɯ ɩɚɪɚɡɢɬɨɜ ɥɨɤɚɥɢɡɭɸɬɫɹ ɜ
ɩɨɥɨɫɬɢ ɛɪɸɲɤɚ ɝɥɚɜɧɵɦ ɨɛɪɚɡɨɦ ɜ ɟɝɨ ɡɚɞɧɟɣ ɱɚɫɬɢ ɚ ɢɧɰɢɫɬɢɪɨɜɚɧɧɵɟ - ɜ ɦɭɫɤɭɥɚɬɭɪɟ
ɝɪɭɞɢɢɛɪɸɲɤɚɧɚɫɬɟɧɤɚɯɩɢɳɟɜɨɞɚɢɥɢɜɠɢɪɨɜɨɦɬɟɥɟɜɡɪɨɫɥɵɯɧɚɫɟɤɨɦɵɯ [4]. Ɍɪɟɦɚɬɨɞɵ
ɜɵɡɵɜɚɸɬ ɩɪɨɫɬɨɝɨɧɢɦɨɡ- ɡɚɛɨɥɟɜɚɧɢɟɜɨɞɧɵɯɢɨɤɨɥɨɜɨɞɧɵɯɩɬɢɰɇɚɱɬɨɭɤɚɡɵɜɚɸɬȾɒ
90
ɘɥɩɦɩɞɣɺ ɡɣɝɩɭɨɶɰ
Ecology of animals
əɞ ɋɩɬɬɣɣ: ɸɥɩɦɩɞɣɺ, ɫɛɢɝɣɭɣɠ. ʌ1 2011
The South of Russia: ecology, development. ʋ
Ʉɭɤɚɲɟɜ Ɋ ɋɉɚɜɥɸɤɌȺɄɪɚɫɧɨɛɵɥɨɜɚȻɎȻɟɥɵɲɟɜ>@. ɉɚɪɚɡɢɬɵɪɚɫɩɪɨɫɬɪɚ
ɧɟɧɵɲɢɪɨɤɨɢɜɟɫɶɦɚɡɧɚɱɢɬɟɥɶɧɨɜɥɢɹɸɬɧɚɪɚɡɥɢɱɧɵɟɫɬɨɪɨɧɵɛɢɨɥɨɝɢɢɢɷɤɨɥɨɝɢɢɫɬɪɟ
ɤɨɡɨɤɚɡɵɜɚɹɧɚɧɢɯɭɝɧɟɬɚɸɳɟɟɜɨɡɞɟɣɫɬɜɢɟ.
ȼɚɠɧɨ ɨɬɦɟɬɢɬɶ ɱɬɨ ɡɚ ɫɱɟɬ ɦɚɫɫɨɜɨɫɬɢ ɢ ɲɢɪɨɤɨɝɨ ɪɚɫɩɪɨɫɬɪɚɧɟɧɢɹ ɫɬɪɟɤɨɡɵ ɹɜɥɹ
ɸɬɫɹɨɞɧɢɦɢɡɫɭɳɟɫɬɜɟɧɧɵɯɡɜɟɧɶɟɜɜɩɢɳɟɜɵɯɜɡɚɢɦɨɨɬɧɨɲɟɧɢɹɯɛɢɨɰɟɧɨɡɨɜ Ɉɧɢ ɨɫɭɳɟɫɬ
ɜɥɹɸɬɩɨɞɞɟɪɠɚɧɢɟɜɢɞɨɜɨɝɨɪɚɜɧɨɜɟɫɢɹɜɫɨɨɛɳɟɫɬɜɚɯɢɫɩɨɥɶɡɭɹɜɤɚɱɟɫɬɜɟɩɢɳɢɦɧɨɝɨɱɢɫ
ɥɟɧɧɵɯɩɪɟɞɫɬɚɜɢɬɟɥɟɣɠɢɜɨɬɧɨɝɨɦɢɪɚ ɉɨɥɶɡɚɤɨɬɨɪɭɸɩɪɢɧɨɫɹɬɷɬɢɧɚɫɟɤɨɦɵɟɜɬɨɦɱɢɫɥɟ
ɢɞɥɹɱɟɥɨɜɟɤɚɡɚɤɥɸɱɚɟɬɫɹɜɢɫɬɪɟɛɥɟɧɢɢɢɦɢɜɛɨɥɶɲɢɯɤɨɥɢɱɟɫɬɜɚɯɦɭɯɤɨɦɚɪɨɜɦɨɲɟɤ
ɫɥɟɩɧɟɣɦɟɥɤɢɯɛɚɛɨɱɟɤ-ɜɪɟɞɢɬɟɥɟɣɫɟɥɶɫɤɨɯɨɡɹɣɫɬɜɟɧɧɵɯɤɭɥɶɬɭɪɅɢɱɢɧɤɢɫɬɪɟɤɨɡɹɜɥɹɸɬ
ɫɹ ɨɞɧɢɦ ɢɡ ɤɨɦɩɨɧɟɧɬɨɜ ɤɨɪɦɨɜɨɣ ɛɚɡɵ ɪɵɛ ɩɬɢɰ ɢ ɛɢɨɢɧɞɢɤɚɬɨɪɚɦɢ ɮɭɧɤɰɢɨɧɢɪɨɜɚɧɢɹ
ɛɢɨɰɟɧɨɡɨɜɢɩɪɨɰɟɫɫɨɜɩɪɨɬɟɤɚɸɳɢɯɜɧɢɯ
ȼɣɜɦɣɩɞɫɛɯɣɲɠɬɥɣɤɬɪɣɬɩɥ
1. ȻɟɥɵɲɟɜȻɎɈɩɪɟɞɟɥɢɬɟɥɶɫɬɪɟɤɨɡɋɢɛɢɪɢɩɨɢɦɚɝɨɢɥɢɱɢɧɨɱɧɵɦɮɚɡɚɦ± ɇɨɜɨɫɢɛɢɪɫɤɇɚɭɤɚ. ± ɫ
2. ȻɟɥɵɲɟɜȻɎɏɚɪɢɬɨɧɨɜȺɘɈɩɪɟɞɟɥɢɬɟɥɶɫɬɪɟɤɨɡɩɨɤɪɵɥɶɹɦ± ɇɨɜɨɫɢɛɢɪɫɤɇɚɭɤɚ± ɫ
3. ȻɟɥɵɲɟɜȻɎɋɬɪɟɤɨɡɵɋɢɛɢɪɢ± ɇɨɜɨɫɢɛɢɪɫɤɇɚɭɤɚ± Ɍɑ± ɫ
4. Ɂɞɭɧȼɂ. ɈɛɢɧɜɚɡɢɢɧɚɫɟɤɨɦɵɯɬɪɟɦɚɬɨɞɚɦɢXIII Ɇɟɠɞɭɧɚɪɨɞɧɵɣɷɧɬɨɦɨɥɨɝɢɱɟɫɤɢɣɤɨɧɝɪɟɫɫ± Ɇɨɫɤɜɚ±9
ɚɜɝɭɫɬɚɝɌɪɭɞɵɆɇɚɭɤɚ±ɌII. ± ɋ±115.
5. ɄɟɬɟɧɱɢɟɜɏȺɏɚɪɢɬɨɧɨɜȺɘ. ɈɩɪɟɞɟɥɢɬɟɥɶɫɬɪɟɤɨɡɄɚɜɤɚɡɚ± ɇɚɥɶɱɢɤɄȻȽɍ± ɫ
6. ɄɪɚɫɧɨɛɵɥɨɜɚɌȺɁɚɪɚɠɟɧɧɨɫɬɶ ɫɬɪɟɤɨɡɅɚɬɜɢɣɫɤɨɣɋɋɊɦɟɬɚɰɟɪɤɚɪɢɹɦɢɬɪɟɦɚɬɨɞɁɨɨɥɨɝɢɱɟɫɤɢɣɠɭɪɧɚɥ ±
1970. ± ɌXLIXȼɵɩ± C. 1290 ± 1297.
7. ɄɭɤɚɲɟɜȾɒɋɬɪɟɤɨɡɵɄɚɡɚɯɫɬɚɧɚ- ɩɪɨɦɟɠɭɬɨɱɧɵɟɯɨɡɹɟɜɚɝɟɥɶɦɢɧɬɨɜɉɪɨɛɥɟɦɵɷɤɨɥɨɝɢɢɝɨɪɧɵɯɪɟɝɢɨ
ɧɨɜ- Ⱦɭɲɚɧɛɟ± ɋ-79.
8. ɉɚɜɥɸɤɊɋ. Ɉɡɚɪɚɠɟɧɧɨɫɬɢɢɦɚɝɢɧɚɥɶɧɨɣ ɫɬɚɞɢɢɫɬɪɟɤɨɡOdonataɩɚɪɚɡɢɬɚɦɢɜ ɭɫɥɨɜɢɹɯɡɚɩɚɞɧɵɯɨɛɥɚɫɬɟɣ
ɍɤɪɚɢɧɵXIII Ɇɟɠɞɭɧɚɪɨɞɧɵɣɷɧɬɨɦɨɥɨɝɢɱɟɫɤɢɣɤɨɧɝɪɟɫɫ± Ɇɨɫɤɜɚ-ɚɜɝɭɫɬɚɝɌɪɭɞɵɆɇɚɭɤɚ±
ɌII. ±ɋ±89
9. ɋɥɭɜɤɨȺȺɑɭɣɤɨɜɘɋɋɭɬɨɱɧɚɹɚɤɬɢɜɧɨɫɬɶɪɚɫɩɪɨɫɬɪɚɧɟɧɢɟɢɫɩɟɤɬɪɵɩɢɬɚɧɢɹɧɟɤɨɬɨɪɵɯɜɢɞɨɜɫɬɪɟɤɨɡ
Ⱥɫɬɪɚɯɚɧɫɤɨɣɨɛɥɚɫɬɢɗɤɨɥɨɝɨ-ɛɢɨɥɨɝɢɱɟɫɤɢɟɩɪɨɛɥɟɦɵȼɨɥɠɫɤɨɝɨɪɟɝɢɨɧɚɢɋɟɜɟɪɧɨɝɨɉɪɢɤɚɫɩɢɹɆɚɬɟɪɢɚɥɵ
ȼɫɟɪɨɫɫɢɣɫɤɨɣɧɚɭɱɧɨ-ɩɪɚɤɬɢɱɟɫɤɨɣɤɨɧɮɟɪɟɧɰɢɢ± ɨɤɬɹɛɪɹ ɝɨɞɚ± Ⱥɫɬɪɚɯɚɧɶɂɡɞ-ɜɨȺɫɬɪɚɯɚɧɫɤɨɝɨɝɨ
ɫɭɞɚɪɫɬɜɟɧɧɨɝɨɩɟɞɚɝɨɝɢɱɟɫɤɨɝɨɭɧɢɜɟɪɫɢɬɟɬɚ± C. 96 ± 98.
10. Chowdhury S. H., Corbet P. S. Feeding-ralated behavior in larva Enallagma cyathigerum (Charpentier) (Zigoptera:
Agrionidae) // Odonatologica, 9ʋ3± 288.
11. Copeland R. S., Okeka W., Corbet P. S Treeholes as larval habitat of the dragon-fly Hadrothemis camarensis (Odonata:
/LEHOOXOLGDHLQ.DNDPHJDIRUHVW.HQ\D9ʋ- P. 129 ± 147.
Bibliography
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1963. ± 387 p.
2. Belyshev B.F., Kharitonov A.Y. Classification Key of the dragonflies by the wings. ± Novosibirsk: the Science, 1977. ± 235 p.
3. Belyshev B.F. Dragonflies of Siberia. ± Novosibirsk: the Science, 1973, Vol.1, Part 1. ± 400 p.
4. Zdun V.I. About the trematode invasion on insects // XIII International Congress of Entomology. ± Moscow, 1968, August 2-9th.Works. M.: the Science, 1971. - Vol. 2. ± 114± 115p.
5. Ketenchiev H.A., Kharitonov A.Y. Determiner of the Dragonflies of the Caucasus. ± Nalchick: KBSU, 1998. ± 118p.
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7. Kukashev D. Sh. Of the Kazakhstan dragonflies are intermediate owners of the parasites. // The Problems of ecology of
the mountain regions. Dushanbe. 1989. ± P. 75 ± 79.
8. Pavlyuk R.S. The infection of adult stage Dragonflies (Odanata) with parasites in the western regions of Ukraine // XIII
International Congress of Entomology. ± Moscow, 1968, August 2-9th. Works. M.: the Science, 1971. ± Vol. 2. ± P. 88± 89.
9. Sluvko A.A., Chuikov Y.S. Diurnal activity, distribution and power spectrum of some species of dragonflies in Astrakhan
region // Ecological and biological problems of the Volga region and the Northern Caspian Sea: Proceedings of the AllRussian scientific-practical conference. 2001 October 1st. ± Astrakhan: Publisher of the Astrakhan State Pedagogical University, 2001. ± P. 96± 98.
10. Chowdhury S. H., Corbet P. S. Feeding-ralated behavior in larva Enallagma cyathigerum (Charpentier) (Zigoptera:
Agrionidae) // Odonatologica, 9ʋ± P. 285 ± 288.
11. Copeland R. S., Okeka W., Corbet P. S Treeholes as larval habitat of the dragon-fly Hadrothemis camarensis (Odonata:
Libellulidae) in Kakamega forest, Kenya, 1996. Vʋ± P. 129 ± 147.
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